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University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln The Handbook: Prevention and Control of Wildlife Wildlife Damage Management, Internet Center for Damage 9-1-1994 Coyotes Jeffrey S. Green Assistant Regional Director USDA-APHIS Animal Damage Control Green, Jeffrey S., "Coyotes" (1994). The Handbook: Prevention and Control of Wildlife Damage. Paper 34. http://digitalcommons.unl.edu/icwdmhandbook/34 This Article is brought to you for free and open access by the Wildlife Damage Management, Internet Center for at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in The Handbook: Prevention and Control of Wildlife Damage by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. For more information, please contact proyster@unl.edu.
Jeffrey S. Green Assistant Regional Director USDA-APHIS- COYOTES Animal Damage Control Lakewood, Colorado 80228 F. Robert Henderson Extension Specialist Animal Damage Control Kansas State University Manhattan, Kansas 66506-1600 Mark D. Collinge State Director USDA-APHIS- Animal Damage Control Boise, Idaho 83705 Fig. 1. Coyote, Canis latrans Damage Prevention and Shed lambing, kidding, and calving Toxicants usually reduce coyote predation. Control Methods M-44 ejector devices for use with Remove carrion to help limit coyote sodium cyanide-loaded plastic Exclusion populations. capsules. They are most effective Produce livestock in confinement. Frightening Agents and during cold weather (fall to spring). Repellents Livestock protection collars (LPC) Herd livestock into pens at night. Guarding dogs: Some dogs have containing Compound 1080 Exclusion fences (net-wire and/or (sodium monofluoroacetate) are electric), properly constructed and significantly reduced coyote predation. registered for use only in certain maintained, can aid significantly in states. reducing predation. Donkeys and llamas: Some are aggressive toward canines and have Fumigants Cultural Methods and Habitat Modification reduced coyote predation. Gas cartridges are registered as a Sonic and visual repellents: Strobe burrow (den) fumigant. Select pastures that have a lower incidence of predation to reduce lights, sirens, propane cannons, and Trapping exposure of livestock to predation. others have reduced predation on both sheep and calves. Leghold traps (Nos. 3 and 4) are Herding of livestock generally reduces effective and are the most versatile predation due to human presence Chemical odor and taste repellents: control tool. during the herding period. None have shown sufficient effectiveness to be registered for Snares are effective where coyotes pass Change lambing, kidding, and calving use. through or under net-wire fences seasons. and in trail sets. PREVENTION AND CONTROL OF WILDLIFE DAMAGE — 1994 Cooperative Extension Division Institute of Agriculture and Natural Resources University of Nebraska - Lincoln United States Department of Agriculture Animal and Plant Health Inspection Service Animal Damage Control C-51 Great Plains Agricultural Council Wildlife Committee
Shooting Aerial hunting is effective in removing Other Methods coyotes where terrain, ground Shooting from the ground is effective. cover, vegetation, regulations, and Denning: Remove adult coyotes and/ Use rabbit distress calls or mimic landownership conditions permit. or their young from dens. howling or other coyote sounds to Hunting with dogs is effective for trail- bring coyotes within shooting ing coyotes from kill sites, locating distance. dens, running coyotes, and assisting with aerial hunting or calling. Identification Habitat weaker lambs are also present. Usu- ally, the stronger lamb is on the In body form and size, the coyote Many references indicate that coyotes periphery and is more active, making (Canis latrans) resembles a small collie were originally found in relatively it more prone to attack than a weaker dog, with erect pointed ears, slender open habitats, particularly the grass- lamb that is at the center of the flock muzzle, and a bushy tail (Fig. 1). lands and sparsely wooded areas of and relatively immobile. Coyotes are predominantly brownish the western United States. Whether or Coyote predation on livestock is gener- gray in color with a light gray to not this was true, coyotes have ally more severe during early spring cream-colored belly. Color varies adapted to and now exist in virtually and summer than in winter for two greatly, however, from nearly black to every type of habitat, arctic to tropic, reasons. First, sheep and cows are usu- red or nearly white in some individu- in North America. Coyotes live in ally under more intensive manage- als and local populations. Most have deserts, swamps, tundra, grasslands, ment during winter, either in feedlots dark or black guard hairs over their brush, dense forests, from below sea or in pastures that are close to human back and tail. In western states, typical level to high mountain ranges, and at activity, thus reducing the opportunity adult males weigh from 25 to 45 all intermediate altitudes. High densi- for coyotes to take livestock. Second, pounds (11 to 16 kg) and females from ties of coyotes also appear in the sub- predators bear young in the spring and 22 to 35 pounds (10 to 14 kg). In the urbs of Los Angeles, Pasadena, raise them through the summer, a pro- East, many coyotes are larger than Phoenix, and other western cities. cess that demands increased nutri- their western counterparts, with males tional input, for both the whelping and averaging about 45 pounds (14 kg) and nursing mother and the growing females about 30 pounds (13 kg). Food Habits young. This increased demand corre- Coyote-dog and coyote-wolf hybrids sponds to the time when young sheep Coyotes often include many items in exist in some areas and may vary or beef calves are on pastures or range- their diet. Rabbits top the list of their greatly from typical coyotes in size, land and are most vulnerable to attack. dietary components. Carrion, rodents, color, and appearance. Also, coyotes in Coyote predation also may increase ungulates (usually fawns), insects the New England states may differ in during fall when young coyotes dis- (such as grasshoppers), as well as live- color from typical western coyotes. perse from their home ranges and stock and poultry, are also consumed. Many are black, and some are reddish. establish new territories. Coyotes readily eat fruits such as These colorations may partially be due watermelons, berries, and other veg- to past hybridization with dogs and etative matter when they are available. General Biology, wolves. True wolves are also present In some areas coyotes feed on human Reproduction, and in some areas of coyote range, particu- refuse at dump sites and take pets larly in Canada, Alaska, Montana, Behavior (cats and small dogs). northern Minnesota, Wisconsin, and Coyotes are opportunistic and gener- Coyotes are most active at night and Michigan. Relatively few wolves ally take prey that is the easiest to during early morning hours (especially remain in the southern United States secure. Among larger wild animals, where human activity occurs), and and Mexico. coyotes tend to kill young, inexperi- during hot summer weather. Where enced animals, as well as old, sick, or there is minimal human interference Range and during cool weather, they may be weakened individuals. With domestic animals, coyotes are capable of catch- active throughout the day. Historically, coyotes were most com- ing and killing healthy, young, and in Coyotes bed in sheltered areas but do mon on the Great Plains of North some instances, adult prey. Prey selec- not generally use dens except when America. They have since extended tion is based on opportunity and a raising young. They may seek shelter their range from Central America to myriad of behavioral cues. Strong, underground during severe weather the Arctic, including all of the United healthy lambs are often taken from a or when closely pursued. Their physi- States (except Hawaii), Canada, and flock by a coyote even though smaller, cal abilities include good eyesight and Mexico. C-52
hearing and a keen sense of smell. feeding and caring for the young. Coy- the site. The quantity of sheep or calf Documented recoveries from severe otes commonly hunt as singles or remains left after a kill vary widely injuries are indicative of coyotes’ pairs; extensive travel is common in depending on how recently the kill physical endurance. Although not as their hunting forays. They will hunt in was made, the size of the animal fleet as greyhound dogs, coyotes have the same area regularly, however, if killed, the weather, and the number been measured at speeds of up to 40 food is plentiful. They occasionally and species of predators that fed on miles per hour (64 km/hr) and can bury food remains for later use. the animal. sustain slower speeds for several miles Pups begin emerging from their den One key in determining whether a (km). by 3 weeks of age, and within 2 sheep or calf was killed by a predator Distemper, hepatitis, parvo virus, and months they follow adults to large is the presence or absence of subcuta- mange (caused by parasitic mites) are prey or carrion. Pups normally are neous (just under the skin) hemor- among the most common coyote dis- weaned by 6 weeks of age and fre- rhage at the point of attack. Bites to a eases. Rabies and tularemia also occur quently are moved to larger quarters dead animal will not produce hemor- and may be transmitted to other ani- such as dense brush patches and/or rhage, but bites to a live animal will. If mals and humans. Coyotes harbor sinkholes along water courses. The enough of the sheep carcass remains, numerous parasites including mites, adults and pups usually remain carefully skin out the neck and head to ticks, fleas, worms, and flukes. Mortal- together until late summer or fall observe tooth punctures and hemor- ity is highest during the first year of when pups become independent. rhage around the punctures. Talon life, and few survive for more than 10 Occasionally pups are found in groups punctures from large birds of prey will to 12 years in the wild. Human activity until the breeding season begins. also cause hemorrhage, but the loca- is often the greatest single cause of tion of these is usually at the top of the Coyotes are successful at surviving coyote mortality. head, neck, or back. This procedure and even flourishing in the presence of becomes less indicative of predation as Coyotes usually breed in February and people because of their adaptable the age of the carcass increases or if the March, producing litters about 9 behavior and social system. They typi- remains are scanty or scattered. weeks (60 to 63 days) later in April and cally display increased reproduction May. Females sometimes breed during and immigration in response to Coyotes, foxes, mountain lions, and the winter following their birth, par- human-induced population reduction. bobcats usually feed on a carcass at the ticularly if food is plentiful. Average flanks or behind the ribs and first con- litter size is 5 to 7 pups, although up to sume the liver, heart, lungs, and other 13 in a litter has been reported. More Damage and Damage viscera. Mountain lions often cover a than one litter may be found in a single Identification carcass with debris after feeding on it. den; at times these may be from Bears generally prefer meat to viscera females mated to a single male. As Coyotes can cause damage to a variety and often eat first the udder from lac- noted earlier, coyotes are capable of of resources, including livestock, poul- tating ewes. Eagles skin out carcasses hybridizing with dogs and wolves, but try, and crops such as watermelons. on larger animals and leave much of reproductive dysynchrony and They sometimes prey on pets and are a the skeleton intact. With smaller ani- behaviors generally make it unlikely. threat to public health and safety when mals such as lambs, eagles may bite off Hybrids are fertile, although their they frequent airport runways and and swallow the ribs. Feathers and breeding seasons do not usually corre- residential areas, and act as carriers of “whitewash” (droppings) are usually spond to those of coyotes. rabies. Usually, the primary concern present where an eagle has fed. regarding coyotes is predation on live- Coyote dens are found in steep banks, Coyotes may kill more than one ani- stock, mainly sheep and lambs. Preda- rock crevices, sinkholes, and under- mal in a single episode, but often will tion will be the focus of the following brush, as well as in open areas. Usu- only feed on one of the animals. discussion. ally their dens are in areas selected for Coyotes typically attack sheep at the protective concealment. Den sites are Since coyotes frequently scavenge on throat, but young or inexperienced typically located less than a mile (km) livestock carcasses, the mere presence coyotes may attack any part of the from water, but may occasionally be of coyote tracks or droppings near a body. Coyotes usually kill calves by much farther away. Coyotes will often carcass is not sufficient evidence that eating into the anus or abdominal area. dig out and enlarge holes dug by predation has taken place. Other evi- Dogs generally do not kill sheep or smaller burrowing animals. Dens vary dence around the site and on the car- calves for food and are relatively from a few feet (1 m) to 50 feet (15 m) cass must be carefully examined to aid indiscriminate in how and where they and may have several openings. in determining the cause of death. attack. Sometimes, however, it is Signs of a struggle may be evident. Both adult male and female coyotes difficult to differentiate between dog These may include scrapes or drag hunt and bring food to their young for and coyote kills without also looking marks on the ground, broken vegeta- several weeks. Other adults associated at other sign, such as size of tracks tion, or blood in various places around with the denning pair may also help in (Fig. 2) and spacing and size of canine C-53
tooth punctures. Coyote tracks tend to be more oval-shaped and compact than those of common dogs. Nail marks are less prominent and the tracks tend to follow a straight line more closely than those of dogs. The average coyote’s stride at a trot is 16 to 18 inches (41 to 46 cm), which is typi- cally longer than that of a dog of simi- lar size and weight. Generally, dogs 3" attack and rip the flanks, hind quar- ters, and head, and may chew ears. The sheep are sometimes still alive but 5" Coyote may be severely wounded. Accurately determining whether or not predation occurred and, if so, by what species, requires a considerable amount of knowledge and experience. Evidence must be gathered, pieced Wolf together, and then evaluated in light of the predators that are in the area, the time of day, the season of the year, and numerous other factors. Sometimes even experts are unable to confirm the cause of death, and it may be neces- sary to rely on circumstantial informa- tion. For more information on this subject, refer to the section Procedures for Evaluating Predation on Livestock 2 1/2" 4" and Wildlife, in this book. Red fox Legal Status Large dog The status of coyotes varies depending Fig. 2. Footprints of canid predators on state and local laws. In some states, including most western states, coyotes are classified as predators and can be Federal statutes that pertain to wildlife Damage Prevention and taken throughout the year whether or damage control include the Federal not they are causing damage to live- Insecticide, Fungicide, and Rodenti- Control Methods stock. In other states, coyotes may be cide Act (FIFRA), which deals with For managing coyote damage, a vari- taken only during specific seasons and using toxicants, and the Airborne ety of control methods must be avail- often only by specific methods, such as Hunting Act, which regulates aerial able since no single method is effective trapping. Night shooting with a spot- hunting. in every situation. Success usually light is usually illegal. Some state laws Laws regulating coyote control are not involves an integrated approach, com- allow only state or federal agents to necessarily uniform among states or bining good husbandry practices with use certain methods (such as snares) to even among counties within a state, effective control methods for short take coyotes. Some states have a provi- and they may change frequently. A periods of time. Regardless of the sion for allowing the taking of pro- 1989 Supreme Court action established means used to stop damage, the focus tected coyotes (usually by special that it was not legal to circumvent the should be on damage prevention and permit) when it has been documented laws relative to killing predators, even control rather than elimination of coy- that they are preying on livestock. In to protect personal property (livestock) otes. It is neither wise nor practical to some instances producers can apply from predation. kill all coyotes. It is important to try to control methods, and in others, control prevent coyotes from killing calves or must be managed by a federal or state sheep for the first time. Once a coyote agent. Some eastern states consider the has killed livestock, it will probably coyote a game animal, a furbearer, or a continue to do so if given the protected species. C-54
opportunity. Equally important is taking action as quickly as possible to stop coyotes from killing after they start. 1/4" x 4 1/2" carriage bolt Exclusion galvanized woven wire Most coyotes readily cross over, under, or through conventional live- stock fences. A coyote’s response to a fence is influenced by various factors, 72" galvanized 3" treated pole including the coyote’s experience and 7' long fencing motivation for crossing the fence. Total 6" stays 69" exclusion of all coyotes by fencing, 4" vertical especially from large areas, is highly spacing at top 1 1/2" at the unlikely since some eventually learn to 56" bottom either dig deeper or climb higher to defeat a fence. Good fences, however, can be important in reducing preda- tion, as well as increasing the effective- ness of other damage control methods (such as snares, traps, or guarding 23" apron animals). (old fencing) Recent developments in fencing equip- ment and design have made this technique an effective and economi- cally practical method for protecting 28" sheep from predation under some grazing conditions. Exclusion fencing may be impractical in western range sheep ranching operations. Fig. 3. Barrier fence with wire overhang and Net-Wire Fencing. Net fences in buried apron. good repair will deter many coyotes from entering a pasture. Horizontal spacing of the mesh should be less than 6 inches (15 cm), and vertical Electric Fencing. Electric fencing, The number of spacings between wires spacing less than 4 inches (10 cm). Dig- used for years to manage livestock, has varies considerably. A fence of 13 ging under a fence can be discouraged recently been revolutionized by the strands gave complete protection to by placing a barbed wire at ground introduction of new energizers and sheep from coyote predation in tests at level or using a buried wire apron new fence designs from Australia and the USDA’s US Sheep Experiment Sta- (often an expensive option). The fence New Zealand. The chargers, now also tion (Fig. 4). Other designs of fewer should be about 5 1/2 feet (1.6 m) high manufactured in the United States, wires were effective in some studies, to discourage coyotes from jumping have high output with low impedance, ineffective in others. over it. Climbing can usually be pre- are resistant to grounding, present a The amount of labor and installation vented by adding a charged wire at minimal fire hazard, and are generally techniques required vary with each the top of the fence or installing a wire safe for livestock and humans. The type of fencing. High-tensile wire overhang. fences are usually constructed of fences require adequate bracing at cor- smooth, high-tensile wire stretched to Barrier fences with wire overhangs ners and over long spans. Electric fenc- a tension of 200 to 300 pounds (90 to and buried wire aprons were tested in ing is easiest to install on flat, even 135 kg). The original design of electric Oregon and found effective in keeping terrain. Labor to install a high-tensile fences for controlling predation con- coyotes out of sheep pastures (Fig. 3). electric fence may be 40% to 50% less sisted of multiple, alternately charged The construction and materials for than for a conventional livestock fence. and grounded wires, with a charged such fencing are usually expensive. trip wire installed just above ground Labor to keep electric fencing func- Therefore, fences of this type are rarely level about 8 inches (20 cm) outside the tional can be significant. Tension of the used except around corrals, feedlots, main fence to discourage digging. wires must be maintained, excessive or areas of temporary sheep confine- Many recent designs have every wire vegetation under the fence must be ment. charged. removed to prevent grounding, dam- C-55
age from livestock and wildlife must Fiberglass line post be repaired, and the charger must be checked regularly to ensure that it is operational. Coyotes and other predators occasion- 66' ally become “trapped” inside electric fences. These animals receive a shock Fiberglass 6' 22' stays as they enter the pasture and subse- 11' quently avoid approaching the fence to escape. In some instances the captured predator may be easy to spot and Fiberglass remove from the pasture, but in line post others, particularly in large pastures with rough terrain, the animal may be Charged wire difficult to remove. + 8" Ground wire - 8" Electric Modification of Existing + 8" Fiberglass Fences. The cost to completely stay - 8" replace old fences with new ones, + 8" whether conventional or electric, can - 8" Trip wire be substantial. In instances where + 8" existing fencing is in reasonably good - 8" 8" condition, the addition of one to sev- + 4" 6" - 4" Note: eral charged wires can significantly + 4" Drawing not to scale enhance the predator-deterring ability - 4" of the fence and its effectiveness for controlling livestock (Fig. 5). A Ground charged trip wire placed 6 to 8 inches level (15 to 230 cm) above the ground about 8 to 10 inches (20 to 25 cm) outside the Fig. 4. High-tensile, electric, antipredator fence. fence is often effective in preventing coyotes from digging and crawling un- der. This single addition to an existing fence is often the most effective and economical way to fortify a fence against coyote passage. If coyotes are climbing or jumping a fence, charged wires can be added to the top and at various intervals. These wires should be offset outside the fence. Fencing companies offer offset brackets to make installation relatively simple. The number of additional wires depends on the design of the original fence and the predicted habits of the predators. Portable Electric Fencing. The Outrigger post advent of safe, high-energy chargers with four wires has led to the development of a variety of portable electric fences. Most are constructed with thin strands of wire running through polyethylene twine or Fig. 5. Existing woven-wire livestock fence modified with electrified wire. ribbon, commonly called polywire or polytape. The polywire is available in single and multiple wire rolls or as mesh fencing of various heights. It can be quickly and easily installed to serve C-56
as a temporary corral or to partition As a final note, fences can pose prob- Counting sheep and calves regularly is off pastures for controlled grazing. lems for wildlife. Barrier fences in par- important in large pastures or areas ticular exclude not only predators, but with heavy cover where dead livestock Perhaps the biggest advantage of port- also many other wildlife species. This could remain unnoticed. It is not able electric fencing is the ability to set fact should be considered where fenc- unusual for producers who do not up temporary pens to hold livestock at ing intersects migration corridors for regularly count their sheep to suffer night or during other predator control wildlife. Ungulates such as deer may fairly substantial losses before they activities. Portable fencing increases attempt to jump fences, and they occa- realize there is a problem. Determining livestock management options to sionally become entangled in the top with certainty whether losses were due avoid places or periods of high preda- wires. to coyotes or to other causes may tion risk. Range sheep that are not become impossible. accustomed to being fenced, however, Cultural Methods and Habitat may be difficult to contain in a port- Season and Location of Lambing Modification able fence. or Calving. Both season and location At the present time, there are no docu- of lambing and calving can signifi- Fencing and Predation Manage- mented differences in the vulnerability cantly affect the severity of coyote pre- ment. The success of various types of of various breeds of sheep to coyote or dation on sheep or calves. The highest fencing in keeping out predators has dog predation because there has been predation losses of sheep and calves ranged from poor to excellent. Density very little research in this area. Gener- typically occur from late spring and behavior of coyotes, terrain and ally, breeds with stronger flocking through September due to the food vegetative conditions, availability of behaviors are less vulnerable to preda- requirements of coyote pups. In the prey, size of pastures, season of the tors. Midwest and East, some lambing or year, design of the fence, quality of A possible cause of increased coyote calving occurs between October and construction, maintenance, and other predation to beef cattle calves is the December, whereas in most of the factors all interplay in determining increased use of cattle dogs in herding. western states lambing or calving how effective a fence will be. Fencing Cows herded by dogs may not be as occurs between February and May. By is most likely to be cost-effective where willing to defend newborn calves from changing to a fall lambing or calving the potential for predation is high, coyotes as those not accustomed to program, some livestock producers where there is potential for a high herding dogs. have not only been able to diversify stocking rate, or where electric modifi- their marketing program, but have cation of existing fences can be used. Flock or Herd Health. Healthy also avoided having a large number of Fencing can be effective when incorpo- sheep flocks and cow/calf herds have young animals on hand during periods rated with other means of predation higher reproductive rates and lower when coyote predation losses are typi- control. For example, combined use of overall death losses. Coyotes often cally highest. guarding dogs and fencing has prey on smaller lambs. Poor nutrition means weaker or smaller young, with Shortening lambing and calving peri- achieved a greater degree of success a resultant increased potential for pre- ods by using synchronized or group than either method used alone. An dation. Ewes or cows in good condi- breeding may reduce predation by electric fence may help keep a guard- tion through proper nutrition will raise producing a uniform lamb or calf crop, ing dog in and coyotes out of a pas- stronger young that may be less vul- thus reducing exposure of small live- ture. If an occasional coyote does pass nerable to coyote predation. stock to predation. Extra labor and through a fence, the guarding dog can facilities may be necessary, however, keep it away from the livestock and Record Keeping. Good record- when birthing within a concentrated alert the producer by barking. keeping and animal identification sys- period. Some producers practice early Fencing can also be used to concen- tems are invaluable in a livestock weaning and do not allow young to go trate predator activity at specific places operation for several reasons. From the to large pastures, thus reducing the such as gateways, ravines, or other standpoint of coyote predation, chance of coyote losses. This also gives areas where the animals try to gain records help producers identify loss orphaned and weak young a greater access. Traps and snares can often be patterns or trends to provide baseline chance to survive. set at strategic places along a fence to data that will help determine what type and amount of coyote damage The average beef cattle calf production effectively capture predators. Smaller control is economically feasible. is about 78% nationwide. First-calf pastures are easier to keep free from Records also aid in identifying critical heifers need human assistance to give predators than larger ones encompass- problem areas that may require atten- birth to a healthy calf about 40% of the ing several square miles (km2). tion. They may show, for example, that time. Cow/calf producers who aver- Fencing is one of the most beneficial losses to coyotes are high in a particu- age 90% to 95% calf crops generally investments in predator damage con- lar pasture in early summer, thus high- check their first-calf heifers every 2 trol and livestock management where lighting the need for preventive hours during calving. Also, most good practical factors warrant its use. control in that area. producers place first-calf heifers in C-57
small pastures (less than 160 acres [64 Some producers reason that coyotes nerable to coyote predation than those ha]). When all cows are bred to pro- are less likely to kill livestock if there is in closer, more open, and smaller pas- duce calves in a short, discreet (e.g. carrion available. This may be a valid tures. In general, a relatively small, 60-day) period, production typically preventative measure if an adequate open, tightly fenced pasture that can increases and predation losses supply of carrion can be maintained be kept under close surveillance is a decrease. The birth weight of calves far away from livestock. If a coyote good choice for birthing livestock that born to first-calf heifers can be de- becomes habituated to a diet of live- are likely targets of coyotes. Past expe- creased by using calving-ease bulls, stock remains, however, it may turn to rience with predators as well as thus reducing birthing complications killing livestock in the absence of car- weather and disease considerations that often lead to coyote predation. casses. Wherever there is easily acces- should also serve as guides in the sible carrion, coyotes seem to gather selection of birthing pastures. Producers who use lambing sheds or and predation losses are higher. Con- pens for raising sheep and small pas- A factor not completely understood is versely, where carrion is generally not tures or paddocks for raising cattle that, at times, coyotes and other preda- available, losses are lower. A study in have lower predation losses than those tors will kill in one pasture and not in Canada showed that the removal of who lamb or calve in large pastures or another. Therefore, changing pastures livestock carcasses significantly on open range. The more human pres- during times of loss may reduce pre- reduced overwinter coyote popula- ence around sheep, the lower the pre- dation. There may seem to be a rela- tions and shifted coyote distributions dation losses. Confining sheep entirely tionship between size of pasture and out of livestock areas. to buildings virtually eliminates preda- predator losses, with higher loss rates tion losses. Habitat Changes. Habitat features reported in larger pastures. In reality, change in some areas, depending on loss rates may not be related as much Corrals. Although predation can seasonal crop growth. Some cultivated to pasture size as to other local condi- occur at any time, coyotes tend to kill fields are devoid of coyotes during tions such as slope, terrain, and human sheep at night. Confining sheep at winter but provide cover during the populations. Hilly or rugged areas are night is one of the most effective growing season, and a corresponding typically sparsely populated by hu- means of reducing losses to predation. increase in predation on nearby live- mans and are characterized by large Nevertheless, some coyotes and many stock may occur. pastures. These conditions are ideal for dogs are bold enough to enter corrals The creation of nearly 40 million acres coyotes. and kill sheep. A “coyote-proof” corral is a wise investment. Coyotes are more (16 million ha) of Conservation Sheep pastures that contain or are likely to attack sheep in unlighted cor- Reserve Program (CRP) acres may adjacent to streams, creeks, and rivers rals than in corrals with lights. Even if benefit many species of wildlife, tend to have more coyote problems the corral fence is not coyote-proof, the including predators. These acres har- than pastures without such features. mere fact that the sheep are confined bor prey for coyotes and foxes, and an Water courses serve as hunting and reduces the risk of predation. Penning increase in predator populations can travel lanes for coyotes. sheep at night and turning them out at reasonably be predicted. Clearing away weeds and brush from CRP Herders. Using herders with sheep or mid-morning might reduce losses. In areas may reduce predation problems cattle in large pastures can help reduce addition, coyotes tend to be more since predators usually use cover in predation, but there has been a trend active and kill more sheep on foggy or their approach to livestock. Generally, away from herders in recent years rainy days than on sunny days. Keep- the more open the area where live- because of increasing costs and a ing the sheep penned on foggy or rainy stock are kept, the less likely that shortage of competent help. Neverthe- days may be helpful. coyote losses will occur. Often junk less, tended flocks or herds receive Aside from the benefits of livestock piles are located near farmsteads. closer attention than untended live- confinement, there are some problems These serve as good habitat for rabbits stock, particularly in large pastures, associated it. Costs of labor and mate- and other prey and may bring coyotes and problems can be solved before rials associated with building corrals, into close proximity with livestock, they become serious. We recommend herding livestock, and feeding live- increasing the likelihood for opportu- two herders per band of range sheep. stock must be considered. In addition, nistic coyotes to prey on available live- If herders aren’t used, daily or periodic the likelihood of increased parasite stock. Removing junk piles may be a checking of the livestock is a good hus- and disease problems may inhibit good management practice. bandry practice. adoption of confinement as a method of reducing damage. Pasture Selection. If sheep or beef Frightening Devices and cattle are not lambed or calved in Repellents Carrion Removal. Removal and sheds or lots, the choice of birthing proper disposal of dead sheep and pastures should be made with poten- Frightening devices are useful for cattle are important since livestock car- tial coyote predation problems in reducing losses during short periods casses tend to attract coyotes, mind. Lambs and calves in remote or or until predators are removed. The habituating them to feed on livestock. rugged pastures are usually more vul- devices should not be used for long C-58
periods of time when predation is not Some producers use a radio tuned to a problem. To avoid acclimation you an all-night station to temporarily can increase both the degree and dura- deter coyotes, dogs, and other preda- tion of effectiveness by varying the tors. position, appearance, duration, or fre- Vehicles. Parking cars or pickups in quency of the frightening stimuli, or the area where losses are occurring of- using them in various combinations. ten reduces predation temporarily. Many frightening methods have been Effectiveness can be improved or ridiculed in one way or another; never- extended by frequently moving the theless, all of the techniques discussed vehicle to a new location. Some pro- here have helped producers by saving ducers place a replica of a person in livestock and/or buying some time to the vehicle when losses are occurring Fig. 6. Electronic Guard frightening device institute other controls. in the daylight. If predators continue Lights. A study involving 100 Kansas to kill with vehicles in place, the Center developed a frightening device sheep producers showed that using vehicle serves as a comfortable blind in called the Electronic Guard (EG) (Fig. lights above corrals at night had the which to wait and shoot offending 6). The EG consists of a strobe light most marked effect on losses to coy- predators. and siren controlled by a variable otes of all the devices examined. Out Propane Exploders. Propane interval timer that is activated at night of 79 sheep killed by coyotes in corrals, exploders produce loud explosions at with a photoelectric cell. In tests con- only three were killed in corrals with timed intervals when a spark ignites a ducted in fenced pastures, predation lights. Nearly 40% of the producers in measured amount of propane gas. On was reduced by about 89%. The device the study used lights over corrals. most models, the time between explo- is used in Kansas and other states to There was some indication in the sions can vary from about 1 minute to protect cows/calves from coyote pre- study that sheep losses to dogs were 15 minutes. Their effectiveness at dation. Most research on the effective- higher in lighted corrals, but the frightening coyotes is usually only ness of this device, however, has been sample size for dog losses was small temporary, but it can be increased by done on sheep operations. Suggestions and the results inconclusive. Most of moving exploders to different loca- for using the unit differ for pastured the producers (80%) used mercury tions and by varying the intervals be- sheep and range operations. vapor lights that automatically turned tween explosions. In general, the timer on at dusk and off at dawn. To use the EG in fenced pastures (farm on the exploder should be set to fire flocks): Another advantage of lighted corrals is every 8 to 10 minutes, and the location that coyotes are more vulnerable when should be changed every 3 or 4 days. 1. Place EGs above the ground on they enter the lighted area. Coyotes In cattle pastures, these devices should fence posts, trees, or T-posts so they often establish a fairly predictable pat- be placed on rigid stands above the can be heard and seen at greater tern of killing. When this happens in a livestock. Normally, the exploder distances and to prevent livestock lighted corral, it is possible for a pro- should be turned on just before dark from damaging them. ducer to wait above or downwind of and off at daybreak, unless coyotes are 2. Position EGs so that rain water can- the corral and to shoot the coyote as it killing livestock during daylight hours. not enter them and cause a malfunc- enters. Red or blue lights may make Motion sensors are now available and tion. the ambush more successful since coy- likely improve their effectiveness, otes appear to be less frightened by though it is still only temporary. 3. Locate EGs so that light can enter them than by white lights. Exploders are best used to reduce the photocell port or window. If losses until more permanent control or positioned in deep shade, they may Revolving or flashing the lights may preventive measures can be imple- not turn on or off at the desired enhance their effectiveness in frighten- mented. In about 24 coyote depreda- times. ing away predators. There is some tion complaints over a 2-year period in speculation that the old oil lamps used 4. The number of EGs used to protect North Dakota, propane exploders in highway construction repelled sheep in fenced pastures depends were judged to be successful in stop- coyotes, presumably because of their on pasture size, terrain features, and ping or reducing predation losses until flickering effect. the amount and height of vegetation offending coyotes could be removed. in or around the pasture. In general, Bells and Radios. Some sheep pro- “Success time” of the exploders ap- at least two units should be used in ducers place bells on some or all of pears to depend a great deal on how small (20 to 30 acres [8 to 12 ha]), their sheep to discourage predators. well they are tended by the livestock level, short-grass pastures. Three to Where effects have been measured, producer. four units should be used in larger however, no difference in losses was Strobe Lights and Sirens. The (40 to 100 acres [16 to 40 ha]), hilly, detected. USDA’s Denver Wildlife Research tall grass, or wooded pastures. C-59
5. Don’t use EGs in pastures larger compound that, when applied to Aversive conditioning is well docu- than about 100 acres (40 ha) because sheep, would prevent coyotes from mented for averting rodents from food their effective range is limited. The killing them. Tests were conducted sources, but significant problems must device could be useful in larger pas- with various prey species including be overcome before the method can be tures when placed near areas where rabbits, chickens, and sheep. Some used to reduce coyote predation on sheep congregate and bed at night. repellents were applied by dipping tar- sheep. Coyotes must be induced to eat get animals in them, others were sheeplike baits that have been treated 6. EGs should be placed on high spots, sprayed on, and some were applied in with the aversive chemical. The chemi- where kills have been found, at the neck collars or ear tags. cal must cause sufficient discomfort, edge of wooded areas, near or on such as vomiting, to cause coyotes to bedgrounds, or near suspected Coyotes rely heavily on visual cues avoid other baits. Furthermore, the coyote travelways. They should be while stalking, chasing, and killing avoidance must be transferred to live moved to different locations every their prey. Taste and smell are of lesser sheep and must persist long enough 10 to 14 days to reduce the likeli- importance in actually making the kill. without reinforcement for the method hood of coyotes getting used to These factors may in part account for to offer realistic protection to sheep. them. the fact that the repellent compounds were not able to consistently prevent To date, pen and field tests with To use the EG in open range (herded coyotes from killing, although some of aversive conditioning have yielded or range sheep): the repellents were obviously offensive conflicting and inconclusive results. It 1. The number of EGs used will to coyotes and prevented them from does not appear that aversive condi- depend on the number of sheep in consuming the killed prey. Several tioning is effective in reducing preda- the band and the size of the compounds were tested on sheep tion, but additional field tests would bedground. Four units should be under field conditions, but none be useful. used to protect bands of 1,000 ewes appeared to offer significant, pro- Guarding Animals. and their lambs. longed protection. Livestock Guarding Dogs. A live- 2. When possible, place one EG in the If an effective chemical repellent were stock guarding dog is one that gener- center of the bedground and the to be found, the obstacles in bringing it ally stays with sheep or cattle without other three around the edge of the to industry use would be significant. harming them and aggressively repels bedground. Try to place the units The compound would not only need predators. Its protective behaviors are on coyote travelways. to be effective, but also persistent largely instinctive, but proper rearing enough to withstand weathering while plays a part. Breeds most commonly 3. EGs should be placed on high posing no undue risk to the sheep, used today include the Great Pyrenees, points, ridge tops, edges of clear- other animals, or the environment. It Komondor, Anatolian Shepherd, and ings, or on high rocks or would also have to withstand the rig- Akbash Dog (Fig. 7). Other Old World outcroppings. Hang the devices on orous Environmental Protection breeds used to a lesser degree include tree limbs 5 to 7 feet (1.5 to 2.1 m) Agency (EPA) approval process. Maremma, Sharplaninetz, and Kuvasz. above ground level. If used above Crossbreeds are also used. timberline or in treeless areas, hang High-frequency sound has also been them from a tripod of poles. tested as a repellent for coyotes, but The characteristics of each sheep the results were no more encouraging operation will dictate the number of 4. Herders who bed their sheep tightly than for chemical repellents. Coyotes, dogs required for effective protection will have better results than those like dogs, responded to particular from predators. If predators are scarce, who allow sheep to bed over large sound frequencies and showed some one dog is sufficient for most fenced areas. Sheep that are bedded about aversion to sounds broadcast within pasture operations. Range operations 200 yards (166 m) or less in diam- one foot (30 cm) of their ear. Research- often use two dogs per band of sheep. eter, or are spread out not more ers, however, were unable to broad- The performance of individual dogs than 200 to 400 yards (166 to 332 m) cast the sound a sufficient distance to will differ based on age and experi- along a ridge top, can usually be test the effects under field conditions. ence. The size, topography, and habitat protected with EGs. of the pasture or range must also be Aversive Conditioning. The objec- Repellents. The notion of repelling considered. Relatively flat, open areas tive of aversive conditioning is to feed coyotes from sheep or calves is can be adequately covered by one dog. a coyote a preylike bait laced with an appealing, and during the 1970s, uni- When brush, timber, ravines, and hills aversive agent that causes the coyote versity and government researchers are in the pasture, several dogs may be to become ill, resulting in subsequent tested a wide variety of potentially re- required, particularly if the sheep are avoidance of the prey. Most of the pellent chemical compounds on sheep. scattered. Sheep that flock and form a research on this technique has Both olfactory (smell) and gustatory cohesive unit, especially at night, can involved the use of lithium chloride, a (taste) repellents were examined. The be protected by one dog more effec- salt, as the aversive agent. underlying objective was to find a tively than sheep that are continually C-60
working unless the dog is chained or confined while the control takes place. Dogs caught in a steel trap set for predators are rarely injured seriously if they are found and released within a reasonable period of time. If snares and traps are used where dogs are working, the producer should: (1) encourage the use of sets and devices that are likely not to injure the dog if it is caught, and (2) know where traps and snares are set so they can be checked if a dog is missing. Aerial hunting, as well as calling and shoot- ing coyotes, should pose no threat to guarding dogs. Ensuring the safety of the dog is largely the producer’s responsibility. Dogs may be viewed as a first line of Fig. 7. Livestock guarding dog (Akbash dog) defense against predation in sheep and cow/calf operations in some cases. Their effectiveness can be enhanced by scattered and bedded in a number of to the possibility that the dog may good livestock management and by locations. roam onto their property and that eliminating predators with suitable some predator control devices such as removal techniques. The goal with a new puppy is to chan- traps, snares, and M-44s present a nel its natural instincts to produce a Donkeys. Although the research has danger to it. Many counties enforce mature guardian dog with the desired not focused on donkeys as it has on stringent laws regarding owner characteristics. This is best accom- guarding dogs, they are gaining in responsibility for damage done by plished by early and continued asso- popularity as protectors of sheep and roaming dogs. It is in the best interests ciation with sheep to produce a bond goat flocks in the United States. A of the owner, dog, and community to between the dog and sheep. The opti- recent survey showed that in Texas train the dog to stay in its designated mum time to acquire a pup is between alone, over 2,400 of the 11,000 sheep area. 7 and 8 weeks of age. The pup should and goat producers had used donkeys be separated from litter mates and The use of guarding dogs does not as guardians. placed with sheep, preferably lambs, eliminate the need for other predation The terms donkey and burro are syn- in a pen or corral from which it can’t control actions. They should, however, onymous (the Spanish translation of escape. This socialization period be compatible with the dog’s behavior. donkey is burro) and are used inter- should continue with daily checks Toxicants (including some insecticides changeably. Donkeys are generally from the producer until the pup is and rodenticides) used to control vari- docile to people, but they seem to about 16 weeks old. Daily checks don’t ous pest species can be extremely haz- have an inherent dislike of dogs and necessarily include petting the pup. ardous to dogs and are therefore not other canids, including coyotes and The primary bond should be between compatible with the use of guarding foxes. The typical response of a don- the dog and the sheep, not between the dogs. key to an intruding canid may include dog and humans. The owner, how- braying, bared teeth, a running attack, The M-44 is particularly hazardous to ever, should be able to catch and han- kicking, and biting. Most likely it is dogs. Some people have successfully dle the dog to administer health care acting out of aggression toward the trained their dogs to avoid M-44s by or to manage the livestock. At about 4 intruder rather than to protect the allowing the dog to set off an M-44 months, the pup can be released into a sheep. There is little information on a filled with pepper or by rigging the larger pasture to mingle with the other donkey’s effectiveness with noncanid device to a rat trap. The unpleasant sheep. predators such as bears, mountain experience may teach the dog to avoid A guarding dog will likely include M-44s, but the method is not fool- lions, bobcats, or birds of prey. peripheral areas in its patrolling. Some proof—one error by the dog, and the Reported success of donkeys in reduc- have been known to chase vehicles result is usually fatal. With the excep- ing predation is highly variable. and wildlife and threaten children and tion of toxic collars, which are not legal Improper husbandry or rearing prac- cyclists. These activities should be dis- in all states, toxicants should not be tices and unrealistic expectations couraged. Neighbors should be alerted used in areas where guarding dogs are C-61
probably account for many failures. Llamas. Like donkeys, llamas have an Toxicants Donkeys are significantly cheaper to inherent dislike of canids, and a grow- Pesticides have historically been an obtain and care for than guarding ing number of livestock producers are important component in an integrated dogs, and they are probably less prone successfully using llamas to protect approach to controlling coyote dam- to accidental death and premature their sheep. A recent study of 145 age, but their use is extremely mortality than dogs. They may pro- ranches where guard llamas were used restricted today by federal and state vide a longer period of useful life than to protect sheep revealed that average laws. All pesticides used in the United a guarding dog, and they can be used losses of sheep to predators decreased States must be registered with the EPA with relative safety in conjunction with from 26 to 8 per year after llamas were under the provisions of FIFRA and snares, traps, M-44s, and toxic collars. employed. Eighty percent of the must be used in accordance with label ranchers surveyed were “very satis- Researchers and livestock producers directions. Increasingly restrictive fied” or “satisfied” with their llamas. have identified several key points to regulations implemented by EPA Llamas reportedly bond with sheep consider when using a donkey for pre- under the authority of FIFRA, the within hours and offer advantages dation control: National Environmental Policy Act over guarding dogs similar to those (NEPA), presidential order, and the 1. Use only a jenny or a gelded jack. described for donkeys. Endangered Species Act have resulted Intact jacks are too aggressive and Other Animals. USDA’s Agricultural in the near elimination of toxicants may injure livestock. Some jennies Research Service tested the bonding of legally available for predator damage and geldings may also injure live- sheep to cattle as a method of protect- control. stock. Select donkeys from ing sheep from coyote predation. medium-sized stock. The only toxicants currently registered There was clearly some protection for mammalian predator damage con- 2. Use only one donkey per group of afforded the sheep that remained near trol are sodium cyanide, used in the sheep. The exception may be a cattle. Whether this protection resulted M-44 ejector device, and Compound jenny with a foal. When two or from direct action by the cattle or by 1080 (sodium monofluoroacetate), for more adult donkeys are together or the coyotes’ response to a novel stimu- use in the livestock protection collar. with a horse, they usually stay lus is uncertain. Later studies with These toxicants are Restricted Use Pes- together, not necessarily near the goats, sheep, and cattle confirmed that ticides and may be used only by certi- sheep. Also avoid using donkeys in when either goats or sheep remained fied pesticide applicators. Information adjacent pastures since they may near cattle, they were protected from on registration status and availability socialize across the fence and ignore predation by coyotes. Conversely, of these products in individual states the sheep. goats or sheep that grazed apart from may be obtained from the respective cattle, even those that were bonded, 3. Allow about 4 to 6 weeks for a state’s department of agriculture. were readily preyed on by coyotes. naive donkey to bond to the sheep. Sodium Cyanide in the M-44. The Stronger bonding may occur when There are currently no research data M-44 is a spring-activated device used a donkey is raised from birth with available on the ideal ratio of cattle to to expel sodium cyanide into an sheep. sheep, the breeds of cattle, age of cattle animal’s mouth. It is currently regis- most likely to be used successfully, or 4. Avoid feeds or supplements con- tered by EPA for use by trained per- on the size of bonded groups to obtain taining monensin or lasolacid. They sonnel in the control of depredating maximum protection from predation. are poisonous to donkeys. coyotes, foxes, and dogs. Multispecies grazing offers many 5. Remove the donkey during lamb- advantages for optimum utilization of The M-44 consists of a capsule holder ing, particularly if lambing in con- forage, and though additional study wrapped in an absorbent material, an finement, to avoid injuries to lambs and experience is needed, it may also ejector mechanism, a capsule contain- or disruption of the lamb-ewe bond. be a tool for coyote damage control. ing approximately 0.9 grams of a pow- dered sodium cyanide mixture, and a 6. Test a new donkey’s response to Any animal that displays aggressive 5- to 7-inch (15- to 18-cm) hollow stake canids by challenging it with a dog behavior toward intruding coyotes (Fig. 8). For most effective use, set in a pen or small pasture. Discard may offer some benefit in deterring M-44s in locations similar to those for donkeys that don’t show overt predation. Other types of animals good trap sets. Drive the hollow stake aggression to an intruding dog. reportedly used for predation control into the ground. Cock the ejector unit include goats, mules, and ostriches. 7. Use donkeys in smaller (less than and secure it in the stake. Screw the Coyotes in particular are suspicious of 600 acres [240 ha]), relatively open wrapped capsule holder containing the novel stimuli. This behavior is most pastures with not more than 200 to cyanide capsule onto the ejector unit, likely the primary reason that many 300 head of livestock. Large pas- and apply fetid meat bait to the cap- frightening tactics show at least tem- tures with rough terrain and vegeta- sule holder. Coyotes attracted by the porary effectiveness. tion and widely scattered livestock bait will try to bite the baited capsule lessen the effectiveness of a donkey. holder. When the M-44 is pulled, the C-62
their ability to remain effective during The LP collar must be used in conjunc- rain, snow, and freezing conditions. tion with specific sheep and goat hus- c bandry practices to be most effective. While M-44s can be used effectively as Coyote attacks must be directed or tar- part of an integrated damage control geted at collared livestock. This may d program, they do have several disad- be accomplished by temporarily plac- vantages. Because canids are less ing a “target” flock of perhaps 20 to 50 responsive to food-type baits during collared lambs or kids and their warm weather when natural foods are uncollared mothers in a pasture where usually abundant, M-44s are not as coyote predation is likely to occur, effective during warmer months as while removing other sheep or goats they are in cooler weather. M-44s are from that vicinity. In situations where subject to a variety of mechanical mal- LP collars have been used and found functions, but these problems can be ineffective, the common cause of fail- b minimized if a regular maintenance ure has been poor or ineffective target- schedule is followed. A further disad- ing. It is difficult to ensure effective vantage is the tendency for the cyanide targeting if depredations are occurring in the capsules to absorb moisture over infrequently. In most instances, only a time and to cake, becoming ineffective. high and regular frequency of depre- Maximum effectiveness of M-44s is dations will justify spending the time, hampered by the requirement to fol- effort, and money necessary to become low 26 use restrictions established by trained and certified, purchase collars, the EPA in the interest of human and and use them properly. environmental safety. The M-44 is not registered for use in all states, and in The outstanding advantage in using those where it is registered, the state the LP collar is its selectivity in elimi- may impose additional use restric- nating individual coyotes that are tions. A formal training program is responsible for killing livestock. The required before use of M-44s. Some collar may also be useful in removing states allow its use only by federal depredating coyotes that have eluded a ADC specialists, whereas other states other means of control. Disadvantages may allow M-44s to be used by trained include the cost of collars (approxi- and certified livestock producers. mately $20 each) and livestock that must be sacrificed, more intensive Fig. 8. The M-44 device consists of the (a) base, 1080 Livestock Protection Collar. (b) ejector, (c) capsule holder, and (d) cyanide- management practices, and the costs The livestock protection collar (LP containing plastic capsule. and inconvenience of complying with collar or toxic collar) is a relatively use restrictions, including require- new tool used to selectively kill ments for training, certification, and spring-activated plunger propels coyotes that attack sheep or goats. record keeping. One use restriction sodium cyanide into the animal’s Collars are placed on sheep or goats limits the collars to use in fenced pas- mouth, resulting in death within a few that are pastured where coyotes are tures only. They cannot be used to pro- seconds. likely to attack. Each collar contains a tect sheep on open range. Also, collars small quantity (300 mg) of Compound The M-44 is very selective for canids are not widely available, because they 1080 solution. The collars do not because of the attractants used and the are registered for use in only a few attract coyotes, but because of their unique requirement that the device be states. design and position on the throat, triggered by pulling on it. While the most attacking coyotes will puncture use of traps or snares may present a Fumigants the collar and ingest a lethal amount of hazard to livestock, M-44s can be used the toxicant. Unlike sodium cyanide, Carbon monoxide is an effective bur- with relative safety in pastures where 1080 is slow-acting, and a coyote row fumigant recently re-registered by livestock are present. Although not ingesting the toxicant will not exhibit the EPA. Gas cartridges, which contain recommended, they can also be used symptoms or die for several hours. As 65% sodium nitrate and 35% charcoal, in the presence of livestock guarding a result, sheep or goats that are produce carbon monoxide, carbon dogs if the dogs are first successfully attacked are usually killed. The collar dioxide, and other noxious gases when conditioned to avoid the devices. This is registered only for use against ignited. They were registered by the can be done by allowing them to pull coyotes and may be placed only on EPA in 1981 for control of coyotes in an M-44 loaded with pepper. An addi- sheep or goats. dens only. This is the only fumigant tional advantage of M-44s over traps is currently registered for this purpose. C-63
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