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RETURNING GRAZING LAND TO NATURE HELPS MORE THANWOLVES PAGE 4 WILD CANIDS AMONG US: CANWECOEXIST?PAGE 8 PROS AND CONS: THE 2017 MEXICAN WOLF ...
Returning Grazing Land
to Nature Helps More
than Wolves PA G E 4
Wild Canids Among Us:
Can We Coexist? PA G E 8
Pros and Cons: The 2017
Mexican Wolf Recovery
Plan PA G E 1 2
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Pros and Cons:
                                                                         T           he wild Mexican wolf population in
                                                                                     the United States has been grow-
                                                                                     ing, on average, 14 percent annu-
                                                                            ally since 2009. This strong growth proves
                                                                           the inaccuracy of population models from
                                                                          the 2010-2013 recovery team on which
                                                                         I served (with individuals from Michigan
                                                                        Tech University, Turner Endangered Species
                                                                        Fund, the National Park Service and others)
                                                                       and suggests caution in basing conclusions
                                                                      on those models. The 2017 survey detected
                                                                      all-time, record minimum numbers of wolves
                                                                     (114), packs (22), potential breeding pairs (26)
                                                                     and adult Mexican wolves (88) in the wild.
                                                                    Widespread claims of agency mismanagement
                                                                    and genetic crisis—claims made by scientists,
                                                                    media, wildlife associations and members of
                                                                   the public—are being muted by the successful
                                                                  progress of recovery.
                                                                     The 2010-2013 attempt to revise the recov-
                                                                 ery plan was based on what is now decade-old
                                                                information and has been eclipsed by more cur-
                                                               rent data. The 2017 Mexican Wolf Recovery Plan
                                                              is based on analyses led by an independent, inter-
                                                             nationally known endangered species population-

                            Recovering                       viability expert with a group that included some
                                                            former recovery team members. This latest effort
                                                           used a more advanced, customized viability model
                         Mexican Wolves                    with access to an updated pedigree. For more than
                                                          two years, scientists updated all available data to
                                                         determine what is needed for recovery.
                        on a Solid Scientific                The team used wild Mexican wolf data to update:
                                                        effects of inbreeding, mortality rates, catastrophe prob-

                            Foundation                  ability, percent of females breeding, pup production
                                                       and historical range. Previous models were based on
                                                       wolf mortality rates from the northern Rockies, but the
                                                      current plan uses mortality rates from wild Mexican
                        By JIM HEFFELFINGER           wolves in the recovery areas. Previous analyses lacked
                                                     the 15-plus years of data on percent of females breed-
                                                    ing in the wild, considered in the current plan. The last
                                                   recovery team estimated the effects of inbreeding with data
                                                  from only 39 litters, but the current plan is based on 89
                                                 wild Mexican wolf litters from 1998-2014 (50 more litters
                                                 and eight more years of data). Importantly, overall inbreed-
     Jacquelyn Fallon

                                                ing levels of wild-born pups are not increasing—data which
                                                conflicts with claims of a mounting genetic crisis.
                                                                                                  continued on page 14

12   Fall 2018                                                                                     w w w. w o l f . o r g
RETURNING GRAZING LAND TO NATURE HELPS MORE THANWOLVES PAGE 4 WILD CANIDS AMONG US: CANWECOEXIST?PAGE 8 PROS AND CONS: THE 2017 MEXICAN WOLF ...
The 2017 Mexican Wolf
Recovery Plan

                                                                           AdobeStock / RbbrDckyBK
                                   T      he 2017 Mexican wolf (Canis lupus
                                          baileyi) recovery plan is a long
                                          overdue update of the original 1982
                                   plan. It calls for the U.S. Fish and Wildlife
                                   Service (FWS) to establish two genetically
                                   diverse populations in the subspecies’ core
                                   historical range. The southwestern United
                                   States is targeted for a population of ≥ 320
                                   wolves and northern Mexico for a population
                                   of ≥ 200. FWS predicts that 25 to 35 years
            2017 Mexican Wolf      and $260 million will be required to establish
                                   those populations.
                                      Selection of habitat for the population in
              Recovery Plan:       Mexico is not based on the best—or even good—
                                   science, but rather on political pressure. This was
              Really Good on       made clear in the following reaction by Utah to
                                   an early draft of the plan, which indicated that,
                                   because suitable habitat in Mexico was lacking,
             Anti-Wolf Politics,   the recovery region needed to be extended north
                                   to areas outside the subspecies’ historical range:
               Really Bad on       Identification of areas outside the historic range of
                                   the sub-species as part of the recovery area…will be
                                   vigorously opposed (legally and politically) by the Utah
             Pro-Wolf Science      Division of Wildlife Resources and the State of Utah.
                                      Notably, Utah did not indicate that opposition would
                                   be based on scientific grounds. Arizona, New Mexico
                BY MIKE PHILLIPS   and Colorado adopted similar positions.
                                      The dogged press of political considerations by Arizona,
                                   New Mexico, Utah and Colorado ensured that the FWS
                                   would finalize the 2017 plan with undue reliance on
                                                                                continued on page 15

I n t e r n a t i o n a l Wo l f                                                                     Fall 2018 13
RETURNING GRAZING LAND TO NATURE HELPS MORE THANWOLVES PAGE 4 WILD CANIDS AMONG US: CANWECOEXIST?PAGE 8 PROS AND CONS: THE 2017 MEXICAN WOLF ...
Heffelfinger                                      recovery occur mostly outside Mexico.         make unsuitable the historical range
              continued from page 12                            Some advocates with little knowledge          of the Mexican wolf in a relevant time-
                                                                of Mexico contradict the best avail-          frame. Quality wolf habitat exists north
                  The newest plan also takes into               able science and first-hand knowledge         of the Arctic Circle, but we must decide
              account the gradual phase-out of feed-            of Mexican experts. A state-of-the-art        how to restore the historical, ecological
              ing wolves to divert them from livestock          analysis by a binational team identified      role of Mexican wolves. Scientists have
              and includes realistic estimates of con-          28,635 square miles of high quality wolf      recently warned of the perils of pushing
              nectivity between populations. Genetic            habitat in Mexico; clearly Mexico will        recovery north of historical range because
              diversity retention is addressed with             play a vital role in recovery. The same two   of genetic swamping by large wolves of
              objective, measurable and achievable              large recovery areas of suitable habitat in   Canadian origin that disperse from the
              criteria— not ambiguous references to             Mexico were independently identified          Rocky Mountains. (A Yellowstone wolf
              measurements of genetic diversity that            in a jaguar recovery plan. Discounting        already visited Arizona).
              will only lead to endless litigation about        that information would contradict the             We have binational recovery plans
              delisting. To date, human intolerance             Endangered Species Act requirement            for ocelot, jaguar, Sonoran pronghorn,
              has been limiting Mexican wolf recov-             to use best available data in recovery        thick-billed parrot, condor, masked bob-
              ery, not inbreeding depression.                   planning.                                     white, Kemps-Ridley sea turtle and more;
                  Members of the last Mexican wolf                  This updated habitat analysis includes    why shouldn’t the Mexican wolf also
              recovery team asserted that recovery              two measures of human-caused mortal-          benefit from expansion across borders?
              will require three populations of 250             ity (road density and towns). Adding          This recovery plan, based on updated
              Mexican wolves, but this was based on             information on livestock distribution         analyses far more complex and realistic
              theoretical genetic principles, and on the        and protected areas would stack four          than all previous versions, provides for
              outdated, obsolete model from 2010-               redundant layers representing the same        successful Mexican wolf recovery in its
              2013. Despite these shortcomings, it is           issue. Large tracts of private land with      historical range.
              often misrepresented as a threshold for           restricted access in Mexico have the              Efforts are now appropriately focused
              successful recovery. The plan’s founda-           same function as official land designa-       on returning this small wolf subspecies
              tion is an accurate depiction of historical       tions in the U.S. No other carnivore          to its ecological role in the American
              range based on detailed skull and body            recovery plan has a better representa-        Southwest and Mexico. n
              measurements, historical records, genetic         tion of relative distribution of prey on
              differences and measures of ecological            the landscape; past efforts simply used         Supporting Literature
              differentiation.                                  a satellite image of green vegetation as        (with links to full manuscripts)
                  Federal regulations require that              a substitute. Criticism that the analysis
              Mexican wolves be recovered in their              lacks a measure of livestock density is a       Harding, L. E., J. Heffelfinger, D.
              historical range unless it is “unsuitably and     red herring, as no accurate records exist       Paetkau, E. Rubin, J. Dolphin, A.
              irreversibly altered or destroyed.” Earlier       on either side of the border.                   Aoude. 2016. Genetic management
              teams chose to ignore tens of thou-                                                               and setting recovery goals for Mexican
                                                                    Wolves have adapted to environments
                                                                                                                wolves (Canis lupus baileyi) in the wild.
              sands of square miles of suitable habi-           from the Arctic to Arabia, and climate          Biological Conservation 203:151-159.
              tat in Mexico, inappropriately insisting          change is not going to alter, destroy or        https://www.sciencedirect.com/science/
                                                                                                                article/pii/S0006320716304256

                                                                                                                Heffelfinger, J. R., R.M. Nowak, and
                                                                                                                D. Paetkau. 2017. Clarifying historical
                                                                                                                range to aid recovery of the Mexican
                                                                                                                wolf. Journal of Wildlife Management
                                                                                                                81:766-777. https://onlinelibrary.wiley.
                                                                                                                com/doi/full/10.1002/jwmg.21252

                                                                                                                Odell, E.A. Heffelfinger, J.R.
                                                                                                                Rosenstock, S.S., Bishop C.J., Liley, S.,
                                                                                                                González-Bernal, A., Velasco, J.A.,
                                                                                                                Martínez-Meyer, E. 2018. Perils of
                                                                                                                recovering the Mexican wolf outside
                                                                                                                of its historical range. Biological
                                                                                                                Conservation 220:290-298. https://doi.
                                                                                                                org/10.1016/j.biocon.2018.01.020

                                                                                                              Jim Heffelfinger is the Wildlife Science
Larry Lamsa

               At only 25–32 inches tall, the Mexican gray wolf is smaller than its                           Coordinator for the Arizona Game and
               cousin, the gray wolf, with a coat of buff, gray, rust and black.                              Fish Department.

   14         Fall 2018                                                                                                                  w w w. w o l f . o r g
RETURNING GRAZING LAND TO NATURE HELPS MORE THANWOLVES PAGE 4 WILD CANIDS AMONG US: CANWECOEXIST?PAGE 8 PROS AND CONS: THE 2017 MEXICAN WOLF ...
Phillips
continued from page 13
a woefully inadequate habitat-suit-
ability model.
    The model relies on correlation
between climatic and vegetative fac-
tors, and locations where Mexican wolves
were collected historically to identify
suitable habitat for recovery. FWS and
the states justify this reliance by opin-
ing that Mexican wolves evolved to be
precisely adapted to the narrow range
of habitat present within the subspe-
cies’ core historical range in Mexico.
That opinion, however, is undermined
by 1) good science which indicates that
wolves are broadly adaptable to climatic
and vegetative conditions, and 2) the

                                                                                                                                    Jacquelyn Fallon
FWS’s longstanding effort to restore the
subspecies to Arizona and New Mexico
where such conditions differ from those
in Mexico.
    More important, the model is woe-
fully inadequate because of its disregard
for aspects of wolf habitat that good sci-
ence deems essential to recovery: limited    and managing wolves across millions of       failure of science-informed planning
density of livestock, adequate density       acres of private land necessary to sup-      and leadership by FWS simply for the
of wild prey, and large tracts of public     port ≥ 200 animals.                          sake of political expediency.
land where human-caused mortality is             Although the U.S. public supports            Much of the 2017 Mexican wolf
typically low.                               wolf recovery, anti-wolf groups hold         recovery plan is based on the state’s
    Based on the flawed habitat model,       immense political influence in Colorado,     desire to assign to Mexico as much of
the 2017 plan targets 38 percent of          Arizona, New Mexico, and Utah. These         the burden of Mexican wolf recovery as
recovery on an area in Mexico domi-          groups were well served by the scien-        possible—not the best available science.
nated by small tracts of private property    tific gloss the habitat model gives to the   It is worse than a poor replacement for
with abundant livestock and unknown          recovery plan, and by the disastrous         the 1982 plan. Deeply discounting the
numbers of native prey, and where            decision to exclude from it the high-        cardinal role of wolf-livestock interac-
wildlife protection laws are irregularly     quality habitat of the Grand Canyon and      tions and importance of land ownership
enforced and access and safety for field     Southern Rockies ecoregions of north-        ensures that FWS will waste precious
personnel are concerns. The FWS would        ern Arizona/southern Utah and north-         time and millions of dollars, all the while
never target such an area in the U.S. for    ern New Mexico/southern Colorado,            failing to recover Canis lupus baileyi. n
wolf recovery.                               respectively.
    Reliance on the model is already prov-       If politics demanded that FWS            Mike Phillips has served as the executive
ing problematic. Free-ranging Mexican        initially focus on marginal habitat in       director of the Turner Endangered Species
wolves in Mexico are routinely fed artifi-   Mexico by adopting a habitat suitability     Fund and senior advisor to the Turner
cially to promote survival by minimizing     model that discounts the importance          Biodiversity Divisions since he co-founded
                                                                                          both with Ted Turner in 1997. Before that
conflicts with livestock. Such “diver-       of livestock and land ownership, then
                                                                                          Mike worked for the U.S. Fish and
sionary feeding” is required because of      the agency should at least have defined      Wildlife Service and National Park
abundant livestock and relatively scarce     a recovery region that also included         Service leading efforts to restore red
wild prey, suggesting that the area is not   these two ecoregions. Such an approach       wolves to the southeastern U.S. and gray
suitable despite being identified as such    would have facilitated progress once         wolves to the Yellowstone Park. Mike has
by the habitat model. The shortcomings       the inevitable shortcomings of habitat       served in the Montana legislature since
of the model will become even more           in Mexico became undeniable to even          2006, and will hold his Senate seat
apparent as biologists strive to expand      the most ardent opponents to recovery.       through 2020.
recovery in Mexico, completing a record      Failure to advance such a common-
number of initial releases and monitoring    sense approach to recovery represents a

I n t e r n a t i o n a l Wo l f                                                                                       Fall 2018 15
RETURNING GRAZING LAND TO NATURE HELPS MORE THANWOLVES PAGE 4 WILD CANIDS AMONG US: CANWECOEXIST?PAGE 8 PROS AND CONS: THE 2017 MEXICAN WOLF ...
WOLF EXPERTS
    FROM 19
   COUNTRIES
  SHARING 100
 PRESENTATIONS
   Concurrent, Poster,
  Plenary and Keynote

  WOLVES IN A
CHANGING WORLD
OCTOBER 11–14, 2018
MINNEAPOLIS, MN USA
RETURNING GRAZING LAND TO NATURE HELPS MORE THANWOLVES PAGE 4 WILD CANIDS AMONG US: CANWECOEXIST?PAGE 8 PROS AND CONS: THE 2017 MEXICAN WOLF ...
THE LAST GREAT WOLF RESTORATION – COLORADO
                                             A presentation on the concept of reintroducing wolves to Colorado,
            BANQUET                                         focusing on attributes and challenges.
                                                  Attributes • Prey base
            KEYNOTE                             may include: • Amount of public land available
                                                             • Varying eco-regions (high deserts, mountains, etc.)
             MIKE PHILLIPS
                                          Challenges include • Livestock grazing interests/public grazing allotments
                                            factors such as: • Conflicting positions among special-interest
                                                               groups, politicians and USFWS
                                                             • Legislatively sanctioned, nationwide delisting
                                                               of wolves as endangered

 PLENARY SESSIONS

             PANELS                                    Isle Royale                    Red Wolves, Eastern Wolves and
                                             A panel of four will present a             other Canis Mixes in Eastern
         Wolves of the World               summary of ups and downs, and             North America: Taxonomic validity
 Speakers from regions around the                                                        and challenges to recovery
                                         changing conditions affecting wolves
   world, including Asia, Europe,          and trophic systems over 56-plus      A panel of five will discuss topics
  Canada, the Canadian Arctic and           years of research on Isle Royale.   related  to eastern canids, including
 the United States and Mexico, will        They will also address the ways in    implications   for the U.S. Fish and
 cover topics that include progress         which reintroduction of wolves     Wildlife  Service  if science reorganizes
 of recovery in each region, politics      would benefit a future Isle Royale   North American canid species and
 in place to ensure a viable popula-     ecosystem, given the uncertainties of   declares the red wolf synonymous
   tion, issues and problems that         future contributions by ice bridges,    with eastern wolves, or declares
      may need to be addressed.          weather patterns, random population         it a variant of gray wolves.
                                          events, herbivory and other factors
              Ellesmere                    that influence this island system.    SPECIAL PRESENTATION
 A series of speakers will discuss the
                                          Wolf Depredation Control on Livestock    20-Plus Years of Wolves in Yellowstone
 wolves inhabiting Ellesmere Island
and the Canadian Arctic Archipelago,        A panel of experts representing          Doug Smith, project leader for
   focusing on observations at dens      various viewpoints will discuss wolf        the Wolf Restoration Project in
 and other aspects of pack life, and      depredation conflict management.           Yellowstone and Emmy Award
  including a historical summary of          Agencies, field agents, a wolf       winning cinematographer Bob Landis
Dr. L. David Mech’s two-decade study.     advocate and a livestock producer         will present the history of wolves
                                            will discuss key problems and               in Yellowstone since their
          Michipicoten Island              the latest news, and find areas of            reintroduction in 1995.
 An overview of geography, species           agreement and disagreement.
    history, human disturbances                                                                DEBATE
   and recent studies of caribou,
                                                                                       Mexican Wolf Recovery Plan
         wolves and beaver.
                                                                                    A debate between Mike Phillips,
                                                                                   who will discuss and challenge the
                                                                                  current Mexican Wolf Recovery Plan
                                                                                   and Jim deVos, who will defend it.
RETURNING GRAZING LAND TO NATURE HELPS MORE THANWOLVES PAGE 4 WILD CANIDS AMONG US: CANWECOEXIST?PAGE 8 PROS AND CONS: THE 2017 MEXICAN WOLF ...
Gray wolves in Mongolia: changing                Risk effects of wolves on free-ranging
                                      attitudes and current research                 livestock: Can prey-gut microbiome
                                   PRESENTER Uuganbayar Ganbold,                          predict stress response in
                                    biologist and anti-poaching protection                predator–prey interactions?
                                    manager, Hustai Nuruu National Park,                 PRESENTER Azzurra Valerio,
                                                  Mongolia                                  ashington State University,
                                                                                           W
                                                                                             Olympia, Washington
                                   Gray wolves in Estonia: an overview
                                       of population genetics and                Adaptive use of nonlethal strategies for
                                    hybridization with domestic dogs               minimizing wolf–livestock conflict
                                         PRESENTER Liivi Plumer,                     PRESENTER Suzanne Stone, 
                                   Department of Zoology, Institute of Ecology       Northwest Senior Field Representative,
                                    and Earth Sciences, University of Tartu,          Defenders of Wildlife, Boise, Idaho
                                              Harjumaa, Estonia

 SAMPLING OF                       Quantifying the diet of the Alexander
                                                                                      Challenges in wolf management
                                                                                                in Croatia
PRESENTATIONS                        Archipelago wolf in southeast
                                    Alaska using molecular methods
                                                                                         PRESENTER Djuro Huber, 
                                                                                        Faculty of Veterinary Medicine,
                                       PRESENTER Aimee Massey,                      University of Zagreb, Zagreb, Croatia
                                  Oregon State University, Corvallis, Oregon;
                                    Alaska Department of Fish and Game                  The future of wolf poisoning
                                                                                           programs in Canada
                                  Through the eyes of a wolf: quantifying               PRESENTER Hannah Barron,
                                    and classifying the complexities of                      olf Awareness, Inc.,
                                                                                            W
                                         facial signaling in wolves                    Golder, British Columbia, Canada
                                        PRESENTER Elana Hobkirk, 
                                  Durham University, Durham, United Kingdom

AUSTRALIA             BRAZIL               CROATIA                        ESTONIA                     JAPAN

            AUSTRIA            CANADA                    DENMARK                       INDIA                     MONGOLIA

NORWAY                POLAND              SLOVAKIA                         SWEDEN            UNITED KINGDOM
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                                           Comfort and convenience are right on target at Minneapolis Marriott
                                           Northwest. Providing easy access to The Shoppes at Arbor Lakes, this
                                           all-suite hotel in Brooklyn Park is the perfect place to stay during the
                                           symposium. Spread out in upscale accommodations with private
                                           sleeping areas, or wrap up work obligations using ergonomic
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                                           Symposium registrants will receive a special conference rate
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                                           To book your room, go to wolf.org, click on Programs/International
                                           Wolf Symposium/Lodging.
                                           If you prefer to reserve your room over the phone,
                                           call Dana Madich at: 763-536-3332.
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Functional response of wolves to human                                                REGISTRATION
       development across boreal Canada                                  Registration includes 3 breakfasts, 2 lunches, a
           PRESENTER Marco Musiani,                                  reception, all daily break refreshments and materials.
         Department of Biological Sciences,
     Faculty of Science and Faculty of Veterinary                                              Rates go up Sept. 1
           Medicine, University of Calgary,
                                                                        To register or for more information:
              Calgary, Alberta, Canada
                                                                                                Registration fees
           Wolf tracks at the doorstep:
      A 1-year cycle of wolf behavior close
                                                                          International Wolf Center Member. . . . $424
            to houses in Scandinavia                                      After Sept. 1. . . . . . . . . . . . . . . . . . . . . . $450
      PRESENTER Barbara Zimmermann,
                                                                          Non-member. . . . . . . . . . . . . . . . . . . . . $474        > Not a member?
       Scandinavian Wolf Research Project,                               After Sept. 1. . . . . . . . . . . . . . . . . . . . . . $500     Join today at
    Inland Norway University of Applied Sciences,                                                                                           www.wolf.org/support/
                Koppang, Norway                                                                                                             membership/
                                                                           Student registration. . . . . . . . . . . . . . . . $299
         An 18-year spatial and temporal
        analysis of colonizing gray wolves
       (Canis lupus) in disjunct population
                                                           Dietary niche overlap between                      Winter predation patterns of wolves
          PRESENTER Theresa Simpson,                      wolves, coyotes, and hybrids in                        in northwestern Wyoming
          University of Wisconsin-La Crosse,                  a 3-species hybrid zone                          PRESENTER Susannah Woodruff, 
                La Crosse, Wisconsin
                                                              PRESENTER John Benson,                              Regional research coordinator,
                                                             University of Nebraska-Lincoln,                    Alaska Department of Fish and Game
       Shooting wolves: photographs and
                                                                   Lincoln, Nebraska
         the reconfiguration of the wolf
                                                                                                                Humans and their role in shaping
            in nonfiction for children
                                                           Ecology of the Indian gray wolf                      the ecological functions of wolves
         PRESENTER Debra Mitts-Smith,                 (Canis lupus pallipes) in the Suleman                      PRESENTER Thomas Newsome, 
        School of Information Sciences faculty           Range, South Waziristan, Pakistan                     University of Sydney, Sydney, Australia
          member at the University of Illinois
                                                              PRESENTER Abdul Hamid,
                                                          epartment of Wildlife Management,
                                                         D                                                      Challenging the wildlife decision-
     Wolves at Our Door: results of 4-year
                                                      Pir Mehr Ali Shah Arid Agriculture University,                  making infrastructure
     Minnesota education program initiative                       Rawalpindi, Pakistan                            PRESENTER Walter Medwid, 
             PRESENTER Misi Stine, 
                                                                                                           Vermont Wildlife Coalition, Newport, Vermont
      Project Coordinator, Wolves at our Door,            Competition on two legs and four:
              International Wolf Center,              Impacts of wolf-cougar co-occurrence on
               Minneapolis, Minnesota                                                                         Scent-marking and biometeorology:
                                                        resource selection and survival across                An analysis of behavior across canid
                                                             an anthropogenic gradient                          species Gray Wolf (Canis lupus),
    Are livestock-guarding dogs a viable tool
                                                            PRESENTER Lauren Satterfield,                         Red Wolf (Canis rufus), and
     for preventing damages in open-range
                                                     University of Washington, Seattle, Washington                    Coyote (Canis latrans)
      livestock? A case study from Portugal
                                                                                                                    PRESENTER Hannah Jones,
     PRESENTER Francisco Petrucci-Fonseco,
                                                           Individuality in habitat use of
            Groupo Lobo, Lisbon, Portugal                                                                    ardin-Simmons University, Abilene, Texas
                                                                                                             H
                                                         Scandinavian wolves in relation to
                                                            anthropogenic infrastructure
  Patterns of niche partitioning and overlap                                                                  Do novel scavenging opportunities
                                                      PRESENTER David Carricondo-Sanches,                       or risk of interspecific killing by
     between sympatric wolves
AUSTRALIA                        and snow
                              BRAZIL                 InlandCROATIA                      ESTONIA
                                                            Norway University of Applied Sciences,                          JAPAN
                                                                                                            wolves influence occupancy and activity
   leopards in the mountains of central Asia                     Koppang,Norway
         PRESENTER Shannon Kachel, 
                                                                                                                patterns of smaller carnivores?
    University of Washington, Seattle, Washington                                                                   PRESENTER David Keiter, 

                AUSTRIA                          CANADA                   DENMARK                            INDIA University of Nebraska,
                                                                                                                                    MONGOLIA
                                                                                                                     School of Natural Resources

 NORWAY                         POLAND                     SLOVAKIA                        SWEDEN                   UNITED KINGDOM

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