NORTH FACE SURVEY - Phase II: 2015 - Nevis Landscape Partnership

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NORTH FACE SURVEY - Phase II: 2015 - Nevis Landscape Partnership
NORTH FACE SURVEY

                  Phase II: 2015
                  Report compiled by Lewis Pate
    Environmental Project Officer at Nevis Landscape Partnership
                              Jan 2016

The Nevis Partnership is a Charity registered in Scotland, No SCO33418

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NORTH FACE SURVEY - Phase II: 2015 - Nevis Landscape Partnership
Contents

Acknowedgements……………………………………………………………………………………………………………………………………………………..3

Summary…………………………………………………………………………………………………………………………………………………………………….4

Aims & Objectives……………………………………………………………………………………………………………………………………………………….5

Project Background & Description…………………………………………………………………………………………………………………………6,7,8

Training day……………………………………………………………………………………………………………………………………………………………9,10

2015 Survey………………………………………………………………………………………………………………………………………………….…11,12,13

Areas visited, teams and notable features……………………………………………………………………………..……………………14,15,16,17

Recommendations…………………………………………………………………………………………………………………………………………………….18

Appendix I – Iain Strachan, Botanical Report………………………………………………………………………………………..……..19,20,21,22

Appendix II – MVE, Geological Report….……………………………………………………………………………. 23,24,25,26,27,28,29,30,31

Appendix III – Gordon Rothero, Notable Bryophytes…………………………………………………………………….32,33,34,35,36,37,38

Appendix IV – Gordon Rothero, Nationally Rare & Scarce Bryophytes (2014 Survey)…………………………………………39,40,41

Appendix V – Mike Pescod, Access & Safety Report……………………………………………………………………………………………………42

Appendix VI – Rare & Scarce Vascular Plant Species Not Previously Recorded on Ben Nevis (2014 Survey)………….43,44

Appendix VII – SNH Species ID Cards………………………………………………………………………………………………….………..45,46,47,48

Appendix VIII – North Face Survey Team Contact Details…………………………………………………………………………………………..49

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NORTH FACE SURVEY - Phase II: 2015 - Nevis Landscape Partnership
Acknowledgements

The second ecological and geological survey of the north face of Ben Nevis took place between the 10th and 14th of
August 2015. Building on the success from 2014, the Nevis Landscape Partnership in partnership with SNH continued
with a comprehensive and logistically challenging survey of Britain’s highest mountain.

Funding for the project was kindly provided by the Heritage Lottery Fund (HLF) and Scottish Natural Heritage (SNH).

The challenges of the survey would not have been met without the professional input from the following people and
organisations:

Scottish Natural Heritage – Cathy Mayne
Abacus Mountaineering - Mike Pescod and contracted guides
Midland Valley Exploration – Roddy Muir, Jenny Ellis, Hugh Barron
Rio Tinto Alcan - Landowner
John Muir Trust – Ali Austin & Blair Fyffe
Ian Strachan
Gordon Rothero
NLP Trainee Volunteer Ranger Team
Ed Holt
DMM - Dave Gary
Scottish Mountaineering Club

A major objective of the project has been to raise public awareness of our fascinating and fragile mountain
environments and develop working partnerships between professional mountaineers and conservation bodies.

Images are reproduced courtesy of the survey team, NLP staff and volunteers.

Round-leaved Sundew (Drosera rotundifolia)

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NORTH FACE SURVEY - Phase II: 2015 - Nevis Landscape Partnership
Summary
During August 2015, the Nevis Landscape Partnership, in
collaboration with our working partners, continued to build on
the success of the 2014 survey. We undertook a logistically
challenging and multi-disciplinary survey of Britain’s highest
mountain at 1343mt.
Ben Nevis first attracted botanists and geologists over 200 years
ago, and continues to reveal some of its remaining secrets.
The project team, including geologists, botanists and professional
mountaineers, NLP staff and volunteers, made significant
discoveries, recording new geological features and arctic-alpine
plant species.
Until 2014 many of the 600 metre cliffs, gullies and buttresses remained inaccessible to the scientific community.
Aided by nearly 3,000 meters of rope and low-impact mountaineering techniques the team continued to safely record
new data from previously unexplored areas and revisited challenging areas on the north face of Ben Nevis.
The project is developing a detailed dataset of geological and botanical features which will hopefully provide a guide
for future projects elsewhere, to better understand the habitats that exist on Scotland’s mountains.
Effective partnership working between conservation bodies, professional mountain guides, botanists and geologists
has developed to allow everyone involved to add value to their respective professional knowledge. This will improve
the public’s understanding of our fragile mountain environments with better public engagement and knowledge
sharing.
The NLP and its working partners will continue with one final survey in 2016.

Mike Pescod of Abacus Mountain Guides reviewing survey routes at the CIC hut.

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NORTH FACE SURVEY - Phase II: 2015 - Nevis Landscape Partnership
Primary Aims & Objectives

                           Aim                                                       Objective
 Identify the botanical and geological features of           Advise SNH, working partners and the general public of
 previously inaccessible areas of the north face of Ben      the distribution, location and current condition and of
 Nevis SSSi                                                  these features in detail.
 Build relationships between professionals working in        Improve knowledge base for professional climbers,
 conservation and recreation                                 guides, botanists, geologists and recreational users of
                                                             the mountain
 Deliver three technical surveys of 125ha of normally Develop survey skills and working practices as a blueprint
 inaccessible habitat on the north face of Britain’s highest for future surveys on similar terrain elsewhere
 mountain
 Produce a concise professional report with useful and Inform interested parties of the work undertaken and
 accurate information                                        improve “site condition monitoring”, hopefully bringing
                                                             the unfavourable “vascular plant” feature into
                                                             favourable condition
 Gather geological data to build a three dimensional map Develop a detailed understanding of the processes that
 of the Ben Nevis massif                                     formed the mountain and promote this to a wider
                                                             audience via the “Fieldmove” applications software
 Promote an innovative and exciting project to as wide an Raise awareness of the conservation value of Ben Nevis
 audience as possible, raising the profile of Ben Nevis      SSSi and the pressures put upon it by climate change and
                                                             recreational and commercial use

Descending lower section of Ledge Route

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NORTH FACE SURVEY - Phase II: 2015 - Nevis Landscape Partnership
Project Background

The Ben Nevis Site of Special Scientific Interest (SSSI) is an extensive upland site of 9539ha centred on a massif which
stretches for over 12 kilometres from Ben Nevis, eastwards to Stob Coire na Ceannain, and has an altitudinal range
from 30m above sea level in Glen Nevis to 1344m on the summit.
It contains the following listed features:
(Ref: SNH Site Link, http://gateway.snh.gov.uk/sitelink/siteinfo.jsp?pa_code=192)

 Last Visit Date                       Feature Category                        Feature                      Last assessed Condition

 19/07/13                              Coniferous woodland                     Native pinewood              Unfavourable Recovering
 21/09/15                              Other invertebrates                     Fly assemblage               Favourable Maintained
 16/07/15                              Butterflies                             Small mountain       ringlet Favourable Maintained
                                                                               (Erebia epiphron)
 31/08/15                              Bryophytes                              Bryophyte assemblage         Favourable Maintained
 31/05/15                              Birds – assemblage of Breeding bird assemblage                       Favourable Maintained
                                       breeding birds
 30/09/15                              Mosaic                                  Upland assemblage            Favourable Maintained
 18/07/15                              Igneous petrology                       Caledonian igneous           Favourable Maintained
 08/09/15                              Vascular plants                         Vascular plant assemblage    Unfavourable No Change
 13/06/15                              Broad-leaved, mixed and Upland oak woodland                          Favourable maintained
                                       yew woodland

A large cushion of Gouty-moss (Oedipodium griffithianum) with sporophytes on Tower Ridge

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NORTH FACE SURVEY - Phase II: 2015 - Nevis Landscape Partnership
The range in altitude and geology gives rise to a diversity of habitats including native pine and oak woodlands, montane
cliffs, semi-permanent snow beds, upland heaths and grasslands. A
wide variety of vascular and lower plants are present. The altitude
influences the flora and fauna of the site, further influencing the
range of bird and invertebrate species present.

Ben Nevis and the associated peaks of Aonach Mor and Aonach Beag
form a massif that has the highest land in the UK and contains only
three peaks greater than 1200m out with the Cairngorms. The SSSI
supports an assemblage of upland habitats which includes the only
semi-permanent snow beds outside the Cairngorms, and the most
extensive scree snow-beds in the UK.

The mountain provides unique habitats for rare arctic-alpine flora. The importance of this flora is reflected in the Ben
Nevis SSSI feature for vascular plants, currently in “unfavourable” condition. Some of the key locations that are already
known about are at risk from recreational activity but are also potentially further threatened by changing climate.
The geological features of the site are unlikely to fall into unfavourable condition, however the detailed geological
mapping that MVE undertook in 2014/15 has developed our understanding of the processes that have shaped the
mountain.

Image of the Move software interface from MVE, showing the distribution of the geological data collected during the 2015 survey of the North Face. The coloured
discs represent the 3D orientation of geological contacts, joints and faults that were measured using the Fieldmove Clino application. The data have been projected
onto a digital elevation model of the Lochaber area. Contours are taken from the Ordnance Survey 1:25,000 dataset. View looking to the SW with the Halfway
Lochan on the right.

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NORTH FACE SURVEY - Phase II: 2015 - Nevis Landscape Partnership
A significantly greater area of the 125ha (see Map1 page 21) of the north face of Ben Nevis has now been accessed by
botanists and geologists. Due to the sensitivity of the site and the guiding designations covering it (NSA, SSSI & SAC)
no fixed anchors were placed in the rock.
Rather than employing heavy duty industrial access techniques (such as placing bolts in the rock) the survey team
utilised climbing equipment which can be placed without damaging the site. Many of the sites being accessed were
either damp, ‘flush’ areas or along geological contacts. These locations often coincide with large fractures in the rock
and consequently are full of loose, wet rock. Climbers usually avoid such places for obvious reasons.
The team therefore used many of the usual guiding techniques and had to develop some new approaches. The
experience learned from the 2014 survey made for a much more efficient and targeted approach this year.
The team of mountaineers also brought an extensive and detailed knowledge of the mountain to the scientific survey.
Thousands of days guiding clients on the north face of Ben Nevis gave the geologists and botanists insight into the
mountain’s topography as well taking them safely to previously unrecorded areas.

Ian Strachan training guides on species ID                 Susan Nicol assisting with PPE at the summit

 Man and beast working in harmony                           Rope protectors and belay set-up for surveyors

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NORTH FACE SURVEY - Phase II: 2015 - Nevis Landscape Partnership
The Training Day
It was decided to reduce the “training week from 2014” to one day for the 2015 survey. This was to deliver the project
within budget and, as many of the same team were involved from 2014, it was considered an appropriate timeframe
to bring existing members up to speed. In addition it allowed NLP to maintain a comparable level of input from 2014
for the main survey week.
The day started at NLP offices at An Drochaid and everyone quickly set off to the CIC hut from the north face car park
to work on practical sessions covering:

         survey techniques and data collection – RM & JE from MVE

         GPS, checks and calibration with maps and altimeter – MVE

         geological processes, rock types, features and structures – MVE

         montane habitats and indicator species – IS

         identification of arctic-alpine species – IS

         safety equipment and access logistics – MP

         radios and communications – MP

         rope systems and rigging – MP & team

         Field Move Clino App - MVE

Training Day Team August 2015

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NORTH FACE SURVEY - Phase II: 2015 - Nevis Landscape Partnership
Rescue 8 & shunt set-up for surveyors
                                             Fieldmove Clino App. On IPhone 6

Abseil practice on Organ Pipe Wall
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The Survey (10th – 14 August 2015)

During the survey week many new areas
of the north face were surveyed and
recorded for the first time finding many
new colonies of nationally rare and
scarce plant species: in particular
saxifrages, grasses and mousears, with
additional recording of mosses and
liverworts by Gordon Rothero.

Besides the huge scale of the task and the
challenge of multi-disciplinary surveying
in extremely difficult terrain, the survey
team had to contend with complex
logistical challenges and the need to
physically carry huge amounts of rope,
safety equipment and survey items around the mountain each day. At the end of each day the team had to review
any issues, gather data, and develop plans for the following day.

                                                                      Every morning was arranged with a 7am meet at
                                                                      the North Face car park, and an 8:30 briefing at
                                                                      the CIC hut. Survey teams were finalised, with key
                                                                      sites identified and access routes determined to
                                                                      manage safety and communications. Each
                                                                      location was allocated specific team members
                                                                      with botanists and geologists allocated as
                                                                      required. The daily logistics were recorded on a
                                                                      rota board at the CIC hut for the communications
                                                                      manager.

Morning Planning Meeting at CIC Hut

Communications and radio checks were continued throughout the day with regular updates on team locations and
most importantly managing the hazard of rock fall onto survey teams or the general public. Ed Holt managed the radio
communications from the CIC hut during the week. Ed facilitated effective communication between team members
and monitored public access to the main climbing routes with the aid of a telescope.
The survey was conducted with four teams of two mountaineers either working independently or with a
geologist/botanist as a rope of three. Those teams working without expert guidance would carry out a sweep survey
of technical terrain, noting indicator species and recording any finds on the field App.
To ensure accuracy of records, digital photos of species were taken (with GPS data recorded and checked) and then
checked with the relevant expert back at the base camp at the end of each day. If an area proved of interest, a geologist
or botanist would then be taken back to the key areas. This made the best use of resources and a maintained a balance
between macro and micro scale surveying

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The 2015 survey had the added input of the NLP’s seven trainee volunteer rangers (TVR’s), who were deployed to
strategic locations where they assisted the survey team in various ways. Belay points were monitored on the summit,
200m bags of rope were moved up and down ledge route and along the summit to specific locations and members of
the public were informed of activity to avoid the danger of rock fall from the survey teams.

Iain Strachan training guides on species ID

Before entering terrain with a hazard of rock fall the teams would communicate with the base camp officer and other
teams to ensure the public were not exposed to any risk. Signage around the area was also used, as were numerous
communications during the preceding weeks to the survey on walkers and climbers forums, blogs and websites.
The project was promoted widely on Twitter and Facebook and articles were placed via the following websites and
publications:

Mountain Weather Information Service

Grough Magazine

Press & Journal

UK Climbing

Mountaineering Council of Scotland

SNH Nature of Scotland

                                                          Blair Fyffe JMT

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On the Tuesday Mark Steven from BBC Scotland Out Of Doors spent the day with the team and the project received a
good level of reporting broadcast on Sat 19th Sept and repeated on Sun 20th. Positive feedback was received from
several sources on this programme. It is planned to involve BBC Out Of Doors across additional NLP projects going
forward.

Ian Strachan Summit Interview for BBC Out of Doors

Mara Media had two cameramen on the survey for the Wednesday and filmed for a series “The Highlands”. This is due
for broadcast sometime in 2016 and centres on the connection between people and the landscape of northern
Scotland.

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Mon 10th August

 Team                                    Area Visited                            Notable Species/Features
                                                                                 Recorded (target spp. in bold)

 Donald King, Dave Anderson, Ian         Slingsby’s Chimney to base of           Arctic Mouse-ear, Russet Sedge
 Strachan                                Oservatory Buttress from Coire Leis     and Alpine Saw-wort on traverse,
                                         traverse                                Alpine Lady-fern in Zero Gully

 Al Halewood, Dave Buckett, Gordon       Bowl above the Shroud from Ledge        Alpine Speedwell and Arctic
 Rothero                                 Route to Boomer’s Requiem.              Mouse-ear above the Shroud,
                                                                                 Starwort Mouse-ear at top of
                                         Also plateau edge Ledge Route to        Arch Gully
                                         Castle Ridge

 Mike Pescod, Connor Holdsworth,         Minus Three Gully from Coire Leis       Arctic Mouse-ear, Russet Sedge
 Dan Watson (NTS)                        Traverse                                and Three-flowered Rush on
                                                                                 traverse

 Scott Kirkhope, Will Rowland, Roddy     Base Little Brenva Face                 Starwort Mouse-ear and
 Muir                                                                            Sibbaldia

 Jenny Ellis & Blair Fyffe               North Trident Buttress base

 Cathy Mayne & Cathel Hutchison          Base of Carn Dearg Buttress to Castle   Large stands Russet Sedge, also
 (TVR)                                   Ridge                                   Alpine Lady-fern, Alpine Lady-f
                                                                                 ern

 Lewis Pate, Erika Forbes (TVR), Chris   Moving rope up Ledge Route to top
 Scully (TVR), Allan Audsley (TVR)       of Castle Ridge

 Harris-Harris Brooker (TVR) Phoebe      Signs to shoulder near Lochan Meall
 Webster (TVR)                           an t’Suidhe

Tues 11th August

 Team                                    Area Visited                            Notable Species/Features
                                                                                 Recorded

 Mike Pescod, Will Rowland, Ian          Plateau above upper Cascades            Large pop Highland Saxifrage,
 Strachan                                Cascade                                 also Alpine Meadow-grass,
                                                                                 Starwort Mouse-ear and Arctic
                                                                                 Mouse-ear. New UK altitudinal
                                                                                 records for all four, also for
                                                                                 Mountain Sorrel and Scurvy
                                                                                 Grass

 Mike Pescod, Erika Forbes (TVR),        Summit
 Cathel Hutchison (TVR), Phoebe
 Webster (TVR)

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Scott Kirkhope, Dave Anderson,          East flank of Tower Ridge from crest of   Large stand (100+) Hare’s-foot
 Gordon Rothero                          ridge, descend via Coire Leis             Sedge, also Sibbaldia and
                                                                                   Starwort Mouse-ear

 Al Halewood, Connor Holdsworth,         Broad Gully, west flank Tower Ridge       Highland Saxifrage, Alpine Lady-
 Dan Watson (NTS)                                                                  fern, Sibbaldia and Mountain
                                                                                   Speedwell. Alpine Speedwell
                                                                                   and Sibbaldia in Coire Leis.

 Dave Bucket, Ali Austin (JMT), Roddy    Tower Ridge                               Alpine Speedwell, Arctic Mouse-
 Muir                                                                              ear, Alpine Meadow-grass and
                                                                                   Sibbaldia.

 Donald King, Blair Fyffe, Jenny Ellis   Castle Ridge

 Harris Brooker (TVR), Allan Audsley     Half way lochan/traverse path - safety
 (TVR),

 Cathy Mayne, Chris Scully (TVR)         Coire Leis                                Alpine Lady-fern, Russet Sedge,
                                                                                   Sibbaldia and Northern Buckler-
                                                                                   fern

 Lewis Pate                              Ledge Route and summit with BBC
                                         Scotland Mark Steven Out of Doors

Wed 12th August

 Team                                    Area Visited                              Notable Species/Features
                                                                                   Recorded

 Al Halewood, Dave Anderson, Ian         West of Tower Gully, East flank of        Sibbaldia, also Alpine Speedwell
 Strachan                                Tower Ridge from summit plateau           on Ledge Route

 Conner Holdsworth, Erika Forbes         Coire Leis lochan and abseil posts path   Curved Woodrush (new for Ben
 (TVR), Gordon Rothero                                                             Nevis), also Alpine Meadow-
                                                                                   grass, Starwort Mouse-ear,
                                                                                   Arctic Mouse-ear, Alpine Lady-
                                                                                   fern & Russet Sedge.

 Scott Kirkhope, Will Rowland, Jenny     North Trident Buttress & base mud         Sibbaldia
 Ellis                                   rocks

 Donald King, Dan Watson                 Under the Castle, Moat & Serpent          Starwort Mouse-ear, Alpine
                                                                                   Lady-fern, also Juniper and
                                                                                   Mountain Speedwell

 Dave Bucket & Roddy Muir                First platform WE Buttress                Arctic Mouse-ear

 Mike Pescod & Blair Fyffe               Castle Ridge – rope retrieval

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Cathy Mayne, Ali Austin, Lewis Pate       Ledge Route Easy to summit – rock         Starwort Mouse-ear, Alpine
 & TVR’s                                   site survey on summit                     Speedwell & Sibbaldia en route

                                           Rope to top of Castle Ridge

 Cathel Hutchison (TVR) and Phoebe         Traverse path under Castle Ridge –
 Webster (TVR)                             safety

 Film Crew from Mara Media                 Tbc

Thurs 13th August

 Team                                      Area Visited                              Notable Species/Features
                                                                                     Recorded

 Scott Kirkhope, Connor Holdsworth         No. 3 Gully Buttress from top             Large pops of Starwort Mouse-
 & Ian Strachan                                                                      ear and Highland Saxifrage, also
                                                                                     Arctic Mouse-ear

 Al Halewood, Blair Fyffe & Gordon         South Trident Buttress & No. 4 Gully      Starwort Mouse-ear
 Rothero

 Donald King & Cathy Mayne                 Long Climb to bottom of Basin – Orion     Alpine Lady-fern
                                           Face

 Dave Bucket, Dave Anderson &              Observatory Ridge – descend Ledge
 Roddy Muir                                Route

 Mike Pescod, Ali Austin & Hugh            Secondary Tower Ridge from Italian        Alpine Lady-fern, Sibbaldia.
 Barron (MVE)                              Climb to Pinnacle Buttress, No. 3 & No.   (Drooping and Tufted Saxifrage
                                           4 Gully, descend down Ledge Route         and other known scarce spp also
                                                                                     recorded at top of No 4.)

 Will Rowland & Jim McIntosh (BSBI)        Coire Leis Abseil Posts path and          Good stands of Curved
                                           Buttress east of Bob Run                  Woodrush, Arctic Mouse-ear,
                                                                                     Alpine Speedwell and Sibbaldia,
                                                                                     also Alpine Meadow-grass and
                                                                                     Russet Sedge.

 Susan Nicol, Chris Scully (TVR), Harris   Coire na Ciste – removing rope
 Brooker (TVR)

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Fri 14th August

 Team                                 Area Visited                            Notable Species/Features
                                                                              Recorded

 Mike Pescod, Dave Anderson &         Raeburn’s Buttress Compression          Starwort & Arctic Mouse-ear,
 Cathy Mayne                          Cracks – returned out of South Castle   Alpine Lady-fern, Alpine Saw-
                                      Gully                                   wort, Northern Buckler-fern

                                                                              Cerastium arcticum, Cerastium
                                                                              cerastoides

 Donald Kind & Roddy Muir             Tbc

 Allan Audsley (TVR) & Alex Hogarth   Descend to top car park from CIC with
 (TVR)                                rope

 Dave Bucket, Blair Fyffe & Hugh      Ledge Route to west – descend down      Starwort Mouse-ear, Sibbaldia
 Barron                               No. 5 Gully Top Bowl

 Scot Kirkhope, Gordon Rothero &      Slopes above Raeburn’s Buttress         Starwort & Arctic Mouse-ear,
 Conner Holdsworth                                                            Sibbaldia, Highland Saxifrage

 Al Halewood, Will Rowland & Ian      Indicator Wall RH from summit top       no target spp but new UK alt
 Strachan, Lewis Pate                 70m and descend via Coire Leis          record for Stiff Sedge

 Susan Nicol & Chris Scully (TVR),    Summit
 Harris Brooker (TVR) & Cathel
 Hutchison (TVR)

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Recommendations for 2016:

      Include at least one training day. This should be paid and will include carrying equipment to the CIC Hut, botany
       and geology refreshers as well as further coaching for the experts.
      Consider an additional “open” training day for NGO land staff from other conservation bodies (NTS/JMT/SNH)
       this to be supported by additional paid work with input by the geologists, botanists and guides

      Earlier training of TVR’s on rough ground access and site specific knowledge to reduce management and
       labour-intensity. MP and NLP to assess ability prior to training day.

      Additional independent transport for TVR’s to reduce NLP staff time

      Using the same experts and guides as far as is practical.
      Maintain the current ratio of surveyors/guides.
      Ensure that media teams have their own guides and that they are aware of what they can achieve so as not to
       disrupt the survey too much. Further define media parameters prior to any survey involvement to ensure
       appropriate & relevant promotion of NLP and working partners.

      Use logistical, ecological and geological knowledge of people such as staff from NTS, JMT and NLP – this worked
       well on the 2015 survey.
      Maintain an experienced and paid communications manager at the CIC hut.
      Seek additional sponsored equipment such as waterproof jackets for survey team

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Appendix I
Ben Nevis North Face Survey, year 2, Aug 2015: botanical report

Ian Strachan, 17 Nov 2015

Introduction

The botanical survey in 2015 was again very successful, helped by having the same professional team of climbers
involved as last year. Botanists Ian Strachan and Gordon Rothero were joined for four days (including the training day)
by Dan Watson, NTS Ecologist for Ben Lawers and Glencoe, and for one day by Jim McIntosh, BSBI Scottish Officer.
Both are very experienced mountain botanists and made valuable contributions. Cathy Mayne (SNH), Alison Austin
(JMT) and other team members also contributed botanical skills. The climbers’ familiarity with the North Face was
invaluable, not only for access, but also for identifying potential survey areas which often proved to be of high botanical
interest, such as No 3 Gully buttress, Tower Ridge and the Upper Cascades.

The weather was mixed but did not really hinder survey, and Thursday was a particularly fine day. Late snow-lie limited
access in places, but more of an issue was the very late season, with some species well behind their usual flowering
period, making identification more difficult.

The focus of vascular plant recording was again on Nationally Rare (NR) and Nationally Scarce (NS) species, forming
part of the Ben Nevis SSSI vascular plant assemblage feature (Table 1). Other notable species were also recorded when
seen e.g. Alpine Saw-wort Saussurea alpina and Moss Campion Silene acaulis. We aimed to visit localities not reached
in 2014, notably Tower Ridge, the north side of Carn Dearg Buttress, Coire Leis, Castle Ridge north, Trident Buttress
and several other routes. A mixture of climbing, scrambling and abseiling was used, including short abseils from the
plateau for e.g. Upper Cascades and Indicator Wall (from the summit cairn!). Table X lists the teams and routes covered
each day, together with an indication of key finds (more details below).

On the training day, IS, DW and CM carried out brief ‘refresher’ identification work with the climbers. Laminated ID
cards were available as last year. Most records were made on iPhones using the Midland Valley Fieldmove app, which
had been adapted by the geologists working with the team to record both geological and botanical information. The
botanists also used GPS and notebooks (see next section). Notes were sometimes made of counts or
population/habitat extent, also associated photographs were taken and labelled. Climbing teams surveying without
an expert botanist took photographs to confirm plant identifications.

Data analysis

Finds were recorded in the field in two ways (a) using the Fieldmove app on iPhones, (b) with GPS and notebook. The
latter method was used by IS, DW and CM, although some of their finds were recorded on iPhones instead or as well.
Data from iPhones were downloaded at the end of each day. Notebook records were entered into XL by the recorders
after the survey.

For each survey day, every team that made any plant records has been given a code, from A1 to F4, with a letter for
the day and a number for the team. This was linked to each record for analysis purposes. IPhone data were collated
into a series of spreadsheets for each day/team. The data included a mixture of location records and images
(photographs). The photos were all examined and species identification checked. The botanical site and image records
were combined and sorted by time of day to match photos and locations where possible. The records were imported

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into GIS and the locations converted from lat/long to OS grid reference. The iPhone data was all collated into a single
spreadsheet together with the ‘notebook’ records (which were also imported into GIS) and each record given a unique
number.

Certain difficulties were encountered when ‘cleaning’ the data e.g. photo GRs were usually different from the
apparently corresponding ‘site’ GRs, sometimes very different. Some sites were clearly in the ‘wrong place’, probably
a consequence of the difficulties of using GPS on crags. Judgement had to be applied by examining record details and
locations against maps and air photos in GIS. Decisions made are recorded in the master spreadsheet. Similar problems
were encountered where iPhone and GPS/notebook records coincided. In a few cases the same stands had been
recorded by different teams e.g. in Coire Leis. As far as possible ‘duplicate’ records were identified and omitted from
the final analyses.

Separate layers for each target species were then created in GIS and examined to determine the number of new
‘populations’ in terms of Site Condition Monitoring i.e. stands, or clusters of stands, within a 40m radius. These were
compared with previous records, although this must be provisional in a few cases as the 2014 has still not been fully
collated into GIS.

Findings

Map 1 shows the distribution of all records for target species (i.e. Nationally Rare or Scarce vascular plants) made in
2015. Over 150 target species records were made, most being at new locations. Approximately 68 previously unknown
populations for 10 target species were identified, as detailed in Table 1. A wide range of records were also obtained
for other notable species which will be imported into the BSBI Distributional Database and will contribute to Atlas
2020.

Highlights include:

       A large population of Curved Woodrush (Luzula arcuata) in upper Coire Leis, west of the abseil posts.
        Previously within the SSSI the only recent records were from Aonach Beag, where there is one very large
        population on the plateau and scattered individuals on slopes and gullies. There were old records from the
        Carn Mor Dearg side of Coire Leis which could not be refound in 2015. This new find therefore represents the
        first record for Ben Nevis itself and the second ‘qualifying’ population (>25 plants) for the SSSI. Curved
        Woodrush is classified as ‘Vulnerable’ in the Red Data List.

       A large stand of Hare’s-foot Sedge (Carex lachenalii) with at least 100 flowering shoots on the east flank of
        Tower Ridge, c. 500m SE of the known (smaller) population in Coire na Ciste. Very small populations were
        recorded in 2005 on Aonach Mor and Aonach Beag (the latter could not be found in 2013). This is therefore
        the second ‘qualifying’ population (>25 plants) for the SSSI.

       Large new populations of Highland Saxifrage on the north side of Carn Dearg (Raeburn’s Buttress) and in Coire
        na Ciste (No. 3 Gully Buttress and the Upper Cascades), and of Alpine Speedwell and Arctic Mouse-ear around
        upper Coire Leis.

       More than 10 new populations for each of Alpine lady-fern, Starwort Mouse-ear and Sibbaldia.

       New upper altitudinal limits for Britain & Ireland for several species, including Highland Saxifrage, Alpine
        Meadow-grass, Starwort Mouse-ear, Arctic Mouse-ear, Mountain Sorrel and Scurvy Grass (to be confirmed).

                                                          20
Map 1. Locations of all records for target species made in August 2015 (red squares)

                                                 21
Table 1. Nationally Rare (bold) and Nationally Scarce vascular plant species recorded from new localities on Ben
Nevis North Face, August 2015. * species not recorded previously from Ben Nevis.

                    Common           2015 pops
 Species            name              (records)    Detailed findings

 Athyrium           Alpine            15 (20)      New pops on N face of Castle Ridge, Raeburn’s Buttress, Tower
 distentifolium                                    Ridge (2), Zero Gully, Orion Face, Coire Leis (6). Also several
                    Lady-fern         12 new       known pops re-recorded.

 Carex lachenalii   Hare’s-foot         1 (1)      New population on Tower Ridge E, largest known pop for SSSI
                    Sedge                          (>100 spikes) and a second SCM qualifying population.
                                        new

 Carex saxatilis    Russet            10 (21)      New populations on NE Buttress (1) and Coire Leis (5), also known
                    Sedge                          pops refound/counted below Carn Dearg Buttress (4).
                                       6 new

 Cerastium          Starwort          18 (30)      New pops on Castle Ridge N Face, Raeburn’s Buttress (2),
 cerastoides        Mouse-ear                      Waterfall Gully, Carn Dearg plateau (3), above Trident Buttress,
                                      16 new       No3 Gully Buttress, Upper Cascades, Tower Ridge, Little Brenva
                                                   Face, Coire Leis (4?). Also several known sites relocated

 Cerastium          Arctic            11 (15)      New pops on Raeburn’s Buttress (2), Carn Dearg N Wall, No 3
 nigrescens                                        Gully Buttress (2), Raeburn’s Easy Route, NE Buttress, Upper Coire
                    Mouse-ear          8 new       Leis (large pop). Also two 2014 sites recorded.

 Luzula arcuata*    Curved              1 (8)      Large population in upper Coire Leis, first definite record for Ben
                    Woodrush*                      Nevis and second qualifying pop for SSSI.
                                        new

 Poa alpina         Alpine              5 (8)      Small stands found at several new locations – Raeburn’s Easy
                                                   Route, Douglas Gap, Upper Coire Leis. Also two 2014 sites.
                    Meadow-            3 new
                    grass

 Saxifraga          Highland           7 (11)      Large new pops on Raeburn’s Buttress, No 3 Gully Buttress (2) and
 rivularis          Saxifrage                      Upper Cascades, also new pops on Raeburn’s Easy Route and
                                       6 new       Tower Ridge W. One 2014 site rerecorded.

 Sibbaldia          Sibbaldia         18 (23)      New pops on Raeburn’s Buttress, Waterfall Gully, Tower Ridge E
 procumbens                                        (2), Tower Gully (2), Little Brenva Face (2), upper Coire Leis & rim
                                      11 new       (3). Several 2014 pops also rerecorded.

 Veronica alpina    Alpine              5 (9)      2 large new populations found near S end of Little Brenva Face
                    Speedwell                      and in upper Coire Leis, also new sites on Carn Dearg North Wall
                                       4 new       and Ledge Route.

Note. Poa flexuosa (Wavy Meadow-grass), Saxifraga cernua (Drooping Saxifrage) and Saxifraga cespitosa (Tufted
Saxifrage) were also recorded, but at known sites only.

                                                         22
Appendix II
Summary of Geology for Year 2

Midland Valley Team – Roddy Muir (Monday – Friday)

                        Jenny Ellis (Monday-Wednesday)

                        Hugh Anderson (Thursday-Friday)

Geological Survey – primary objectives

       Use Midland Valley’s Fieldmove Clino application for smartphones to map out the distribution of the
        different rock types in the Allt a ’Mhuilinn and on the North Face;
       Collect data on the spatial orientation of geological contacts, faults and joints;
       Improve our understanding of how the Ben Nevis area has evolved through geological time;
       Construct a 3D model of the Ben Nevis area in Midland Valley’s Move software that can be shared with the
        wider community for teaching and research.

Training Day – Friday 7th August

During the training day we provided a brief overview of our findings from last year’s survey and reviewed the target
areas for the 2015 survey. We also gave a refresher session on the Fieldmove Clino application and how it should be
used for collecting and recording geological and botanical data.

Midland Valley provided five iPhones for the climbing teams and at the end of each day the data was downloaded and
saved before recharging the phones for the following day. We were very impressed with the quality of the data
gathered by the climbing teams during the survey. All of the climbers have made a significant effort to understand
what features the geologists are looking for, and their valuable feedback helped to identify areas that needed to be
investigated in more detail during the survey week. With assistance from Ali Austin from the John Muir Trust we were
able to cover twice as much ground compared with the previous year, and weather permitting, we will be able to visit
the last remaining priority areas during the 2016 survey.

Over the past year, one of us (RM) has also been mapping the outer contact between the Ben Nevis Igneous Complex
and the Dalradian country rocks. Work will continue on mapping the wider Ben Nevis area over the next 12 months in
order to provide sufficient data to build a realistic 3D model of the whole mountain.

                                                         23
Monday 10th August

The contact between the Inner Granite and the Volcanic Pile was traced from the base of the NE Buttress, round the
lower slopes of Little Brenva Face and on up to the crest of the CMD Arête. Snow patches at the head of Coire Leis
made access difficult in places, but the climbers were able to assist RM in reaching all of the relevant rock outcrops.

We have now collected structural data from all exposures of the contact between the Inner Granite and the Volcanic
Pile and we can begin to construct the 3D geometry of this surface in Move.

A preliminary examination of the sedimentary rocks within the Volcanic Pile at the base of North Trident Buttress was
also carried out.

Contact between the Inner Granite (pale pink rock on left of both images) and the Volcanic Pile. Little Brenva Face

Tuesday 11th August

Geological contacts and structural data were mapped in detail during ascents of Tower Ridge and Castle Ridge. In the
gully on the east side of the Douglas Boulder a vertical sheet of andesite approx. 10m wide was observed cutting the
breccias of the Coire na Ciste Formation. This sheet might represent a feeder dyke for the Summit Andesites. RM will
return to this area to map out the sheet in more detail before next year’s survey.

Contact between andesite sheet (grey) and the Coire na Ciste Formation breccias – gully on the east side of the
Douglas Boulder. Image on the RHS shows unweathered andesite with feldspar crystals.

Tower Ridge - Previously unrecorded, fine grained sedimentary rocks similar to those cropping out at the base of North
Trident Buttress were also observed high up on Tower Ridge. The other important discovery on Tower Ridge was the
presence of amygdales within the Summit Andesite Formation. Amygdales are gas bubbles (vesicles) in an igneous
rock that have been filled with a secondary mineral – in this case calcite. The amygdales within the andesite show a

                                                          24
preferred alignment which reflects movement of magma within a lava flow or sill prior to cooling and solidification of
the magma.

Band of fine grained sediments on Tower Ridge – running left to right across the image (orange arrow), just below
the first protection.

Approaching Tower Gap                   Amygdales in the Summit Andesite (to the left of the Krab)

Castle Ridge – Cloud cover hampered some of the investigations on Castle Ridge, but it was still possible to locate the
base of the Summit Andesites overlying the volcanic breccias. A sheet of igneous material (approx. 5m wide) that may
represent a feeder for the Andesites was also observed. Further investigation of Castle Ridge will be required during
next year’s survey.

                                                          25
Feeder dyke (yellow) on the approach to Castle Ridge

                                                       26
Wednesday 12th August

NE Buttress - The orientation of a prominent rhyolite sheet cutting the volcanic breccias on the first platform of NE
Buttress was measured and recorded. Further mapping of the sedimentary rocks forming the lower slopes of Little
Brenva Face has enabled the orientation of the original sedimentary bedding to be determined. Numerous way-up
structures, including small sand volcanoes and water escape structures have been observed and recorded. Previously
unrecorded granite sheet cutting the sedimentary rocks. Further detailed mapping of the different sedimentary units
(fine grained mudstone and coarse conglomerates) making up the Little Brenva Face will be undertaken during next
year’s survey.

                                                 Laminated Mudstone – Little Brenva Face

                                                 Coarse Conglomerate – Little Brenva Face

                                                 Granite Sheet (below red rucsac) cutting sediments – Little Brenva
                                                 Face

North Trident Buttress – the vertical and lateral extent of fine grained sediments overlying the coarse breccias of the
Coire na Ciste Formation was investigated and recorded. Further work will be required next year to look at the internal
structure of the sediments which contain numerous examples of water escape structures. Internal slumping of the
sediments has also been observed.

                                                          27
Fine Grained Sediments(Blue Arrow) – North Trident Buttress. Yellow Arrow = No.5 Gully.

A distinctive andesitic unit with angular fragments of rhyolite crops out in No.5 Gully. A similar unit can be found on
the path 50m upstream from the CIC hut and at the NW margin of the Volcanic Pile beneath Tower Ridge. Detailed
mapping of this unit and the fault rocks in No.5 Gully will be carried out before next year’s survey.

No. 5 Gully and distinctive andesite unit with angular fragments of rhyolite

                                                          28
Thursday 13th August

Observatory Ridge – consists mainly of volcanic breccia with the contact between the overlying Summit Andesites
occurring close to the top of the ridge. Previously unrecorded sheet of granite cutting the volcanic breccia just above
the halfway point.

Observatory Ridge (centre) flanked by the Northeast Buttress to the left and Tower Ridge to the right

Granite sheet (left) cutting volcanic breccia (Observatory Ridge)

Summit – investigated brown weathering igneous rock cropping out between the head of No.4 and No.5 gully. This
unit forms a distinctive scree field on the summit plateau. Appears to from a separate intrusion fed from below by one
or more feeder dykes – further detailed mapping required.

Friday 14th August

                                                          29
Top of Ledge Route and No.5 Gully Bowl – measured and recorded joint sets in the Summit Andesite and examined
one of the feeder dykes for the brown weathering igneous rock on the Summit Plateau.

Distinctive brown weathering igneous rock and feeder dyke in No.5 Gully Bowl.

Photogrammetry

In addition to gathering information on the orientation of geological features, we have also been capturing 3D images
of various rock outcrops to help build the 3D model of the Ben Nevis area. Using digital photographs from smartphones
and more advanced cameras, the images are uploaded to a software package that stitches the 2D images together to
create full 3D images in Move. An example of one of the 3D images is shown below and we hope to be able to generate
a similar model for the whole of the North Face.

3D image (left) of a large outcrop of the Inner Granite – created from 2D photographs (right)

                                                         30
Summary of Geological Data

Image of the Move software interface showing the distribution of the geological data collected during the 2015 survey
of the North Face. The coloured discs represent the 3D orientation of geological contacts, joints and faults that were
measured using the FieldMove Clino application. The data have been projected onto a digital elevation model of the
Lochaber area. Contours are taken from the Ordnance Survey 1:25,000 dataset. View looking to the SW with the
Halfway Lochan on the right.

                                                         31
Appendix III

                                      BEN NEVIS NORTH FACE SURVEY
                                     The notable bryophytes – Gordon Rothero

The main thrust of this survey has been the search for interesting vascular plants and observations of bryophyte
species and communities have been ‘cherry-picked’ during this work. Fortunately the sites that are of interest for
vascular plants are usually good for bryophytes as well but there are some habitats with few vascular plants that have
interesting bryophytes, especially where the snow lies very late into the summer. This is a short account of the more
important bryophyte communities and species; for a much fuller account of the bryophytes on the Ben Nevis SSSI see
Rothero 2005 and 2013. Nomenclature follows Hill et al, 2008.

Where snow lies late into the summer, so-called snowbed communities are widespread on Ben Nevis and the
nationally scarce, chionophilous mosses like Northern Haircap (Polytrichastrum sexangulare, Starke's Fork-moss
(Kiaeria starkei, Sickle-leaved Fork-moss (Kiaeria falcata), Snow Fork-moss (Kiaeria glaciale), Helmet-moss
(Conostomum tetragonum) and Ludwig's Thread-moss (Pohlia ludwigii) are all locally frequent on the more stable
patches of finer gravels in the gullies and along the cornice line. In the latter habitat Kiaeria falcata can form very
large stands (Photo 1) but mostly these species occur in a bryophyte turf of more common species.

                                                          32
Photo 1. A very large stand of Kiaeria falcata just above the top of No. 3 Gully.

There are also important snow-bed liverworts in this habitat including good populations of Alpine Ruffwort (Moerckia
blyttii), Snow Threadwort (Pleurocladula albescens) (Photo 2), Speckled Rustwort (Marsupella sphacelata) and
particularly, Alpine Jagged Notchwort (Lophozia opacifolia). Where the gravels are a bit drier, especially up on the
plateau rim there are scattered stands of Snow Rustwort (Marsupella brevissima), Scarce Silverwort (Anthelia
juratzkana), Book Flapwort (Nardia breidleri) and the nationally rare species Compact Rustwort (Marsupella
condensata) and Pointed Frostwort (Gymnomitrion apiculatum. One other very rare species, Boeck's Rustwort
(Marsupella boeckii), was recorded from wet rocks in this habitat on Ben Nevis, (the old name for the species was
Marsupella nevicensis). I have looked hard for this species in the appropriate habitat during this survey but without
success; it has not been seen on the Ben since 1939.

Photo 2 Pleurocladula albescens in the bryophyte turf in Coire na Ciste

Springs and flushes are also important habitats for bryophytes on the north face as they are for flowering plants,
indeed it is the distinctive pale green colour of the larger patches of the moss Mountain Thread-moss (Pohlia
wahlenbergii var. glacialis) that has often indicated likely target areas for our survey. Other prominent species here
are Pohlia ludwigii and and the much more common Fountain Apple-moss (Philonotis fontana) but mixed with it can
be the nationally scarce Spiral Apple-moss (Philonotis seriata). This mix of the common and the rare is the same with
the liverworts in the springs, the common Water Earwort (Scapania undulata) is usually accompanied by the montane

                                                         33
Marsh Earwort (Scapania uliginosa) and occasionally by the much rarer Floppy Earwort (Scapania paludosa), but they
are quite tricky to separate.

A common moss in the springs and flushes is Claw Brook-moss (Hygrohypnum ochraceum) but in a number of places
this is accompanied by the rare Soft Brook-moss (Hygrohypnum molle). This moss has been known for over 100 years
from the springs at the top of the Red Burn but this survey has produced several new sites; it is now known from No.
5 Gully in at least two places, at the new Hare’s-foot sedge site on the east side of Tower Ridge (Photo 3) and in two
places near the lochan at the head of Coire Leis. In Coire Leis it grows with the rare moss Muehlenbeck's Thread-moss
(Bryum muehlenbeckii) which also occurs in the burn lower down in the coire, and probably elsewhere on the site.
Wet crevices associated with the springs but also on wet crags often have the common Dotted Thyme-moss
(Rhizomnium punctatum) but occasionally there are patches of the larger, montane species Large-leaf Thyme-moss
(Rhizomnium magnifolium).

Photo 3. The habitat for Hygrohypnum molle with Hygrohypnum ochraceum at the Hare’s-foot sedge site on the E
side of Tower Ridge. 2015.

                                                         34
Photo 4 Andreaea nivalis in Coire Leis

Much of the north face is composed of crags of hard acid rocks and these have a limited bryophyte flora and the same
is true of the areas of scree which are usually too mobile for anything other than the most common species. Prominent
here are species of Fringe-moss(Racomitrium) like Bristly Fringe-moss(Racomitrium heterostichum), Slender Fringe-
moss (Racomitrium sudeticum), Woolly Fringe-moss (Racomitrium lanuginosum) and Green Mountain Fringe-moss
(Racomitrium fasciculare) but there are also occasional stands of the much less common Macoun's Fringe-moss
(Racomitrium macounii subsp. alpinum). The granite mosses, species of Andreaea are also very common; Alpine Rock-
moss (Andreaea alpina) is particularly frequent on both solid rock and on gravel, Changeable Rock-moss (Andreaea
mutabilis) occurs in some of the burns and the nationally rare Snow Rock-moss (Andreaea nivalis) is locally abundant
where there is fairly regular irrigation in areas where the snow lies late (Photo 4). On drier rocks in the areas of latest
snow lie there are some shiny, black patches of the very rare Blytt's Rock-moss (Andreaea blyttii), often encrusted with
lichen.

Other uncommon plants that are quite frequent species on the crags include Arctic Fork-moss (Arctoa fulvella), Alpine
Ditrichum (Ditrichum zonatum), Alpine Rustwort (Marsupella alpina) and, where the rock face is often irrigated, the
rosy-red threads of Stabler's Rustwort (Marsupellea stableri). One other species to mention here is the nationally
scarce moss Gouty-moss (Oedipodium griffithianum) which occurs in sheltered crevices over much of the north face;
it is frequent on Tower Ridge, for instance, sometimes in big cushions that look like patches of little lettuces (Photo 5).

                                                            35
Photo 5. A large cushion of Oedipodium griffithianum with sporophytes on Tower Ridge

More base-rich rocks are harder to find and are usually associated with more broken ground, as at the top of No. 4
Gully, and are often associated with stands of the more interesting flowering plants. Occasionally there are calcicole
mosses here like Frizzled Crisp-moss (Tortella tortuosa) and Fine Distichium (Distichium capillaceum) but in general
the species are less demanding. Mougeot's Yoke-moss (Amphidium mougeotii) is quite common and its much rarer
sister species Lapland Yoke-moss (Amphidium lapponicum) occurs in a few places, particularly in the broken ground at
the head of Coire Leis with both Curved woodrush (Luzula arcuata) and Arctic mouse-ear (Cerastium nigrescens). At
the top of No 4 gully there is a good stand of Patent Leskea (Pseudoleskea patens) and this species also occurs on the
broken crags in the basin to the west of Ledge Route where it occurs with Mueller's Silk-moss (Isopterygiopsis
muelleriana).

The more stable areas of scree below the north face are also an important habitat for bryophytes. In the lower screes
there is a patchy development of the internationally important liverwort-rich, oceanic-montane heath community.
The common species here are Taylor's Flapwort (Mylia taylori), Western Earwort (Scapania gracilis), Lesser Whipwort
(Bazzania tricrenata), White Earwort (Diplophyllum albicans), Red Bog-moss (Sphagnum capillifolium) and Woolly
fringe-moss (Racomitrium lanuginosum) and the globally rare and disjunct liverwort Donn's Notchwort
(Anastrophyllum donnianum) is locally frequent. The other characteristic species of this community at this altitude,
Cloud Earwort (Scapania nimbosa) and Birds-foot Earwort (Scapania ornithopodioides) are surprisingly uncommon
given their frequency elsewhere on the Ben Nevis SSSI. Though some of these big liverworts are reasonably common
in the north-west of Scotland, most are very rare in Europe and some have their only European sites here.

                                                         36
Photo 6. Anastrophyllum donnianum amongst broken crags on the west side of Ledge Route

                                                    37
Photo 7. A large patch of Sciuro-hypnum glaciale with Alpine Lady-fern on the east side of Tower Ridge.

Other rare species are associated with crevices in the scree, usually where there is some build up of litter from the
Alpine Lady fern; the most frequent of these are Snow Feather-moss (Sciuro-hypnum glaciale) and Reflexed Feather-
moss (Sciuro-hypnum reflexum), often occurring with Shaded Wood-moss (Hylocomiastrum umbratum) and Downy
Plait-moss (Hypnum callichroum). Both occur with the fern in Coire Leis and Coire na Ciste and Sciuro-hypnum glaciale
has other stands in broken ground on the side of Tower Ridge, at the top of No. 4 Gully and on the west side of Ledge
Route and certainly elsewhere. Other notable species that occur in the interstices of the scree are Muhlenbeck's
Feather-moss (Herzogiella striatella), Greater Pawwort (Barbilophozia lycopodioides), Obtuse Notchwort (Lophozia
obtusa) and Alpine Earwort (Diplophyllum taxifolium).

References

Hill, M.O., Blackstock, T.H., Long, D.G., & Rothero, G.P. 2008. A checklist and census catalogue of British and Irish
bryophytes. Cardiff: The British Bryological Society.

Hill, M.O. and Preston, C.D. 1998. The geographical relationships of British and Irish bryophytes. Journal of Bryology
20: 127-226.

Hodgetts, N.G. 2011. A revised Red List of bryophytes in Britain. Field Bryology 103: 40-49.

Preston, C.D. 2010. A revised list of nationally rare bryophytes. Field Bryology 100: 32-40.

Preston, C.D. 2006. A revised list of nationally scarce bryophytes. Field Bryology 90: 22-29.

Rothero, G. 2005. Ben Nevis SSSI: site condition dossier for bryophytes. Unpublished report for SNH.

Rothero G. P., 2013. Third cycle Site Condition Monitoring report for bryological interest: Ben Nevis SSSI. Scottish
Natural Heritage Commissioned Report

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