Lepidoptera on Forestry Commission Land in England; Conservation Strategy 2007 - 2017 Review 2012/13

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Lepidoptera on Forestry Commission Land in England; Conservation Strategy 2007 - 2017 Review 2012/13
Lepidoptera on Forestry
Commission Land in England;
Conservation Strategy
2007 – 2017
Review 2012/13
Lepidoptera on Forestry Commission Land in England; Conservation Strategy 2007 - 2017 Review 2012/13
Lepidoptera on Forestry Commission Land in England
                Conservation Strategy 2007 – 2017
                         Review 2012/13

This Conservation Strategy is an agreement between Forestry Commission
England and Butterfly Conservation. Forestry Commission England agree to
manage their landholding to encourage and support Lepidoptera and in turn
Butterfly Conservation will monitor species populations and provide
encouragement, management advice and support as required.

Both organisations will strive towards achieving favourable conservation status for
the butterflies and moths identified in this strategy across the priority woodlands.

                    Signed by:

                    Simon Hodgson, Chief Executive,
                    Forestry Commission England

                    Martin Warren, Chief Executive,
                    Butterfly Conservation
Lepidoptera on Forestry Commission Land in England; Conservation Strategy 2007 - 2017 Review 2012/13
Lepidoptera on Forestry Commission Land in England; Conservation Strategy 2007 - 2017 Review 2012/13
Contents
Foreword .................................................................................................................. 4
   Summary ............................................................................................................ 5
1.0       Introduction ................................................................................................... 7
   1.1 Purpose of the Report ................................................................................. 7
2.0       Aims of the Conservation Strategy ................................................................ 8
   2.1 Overall Aim ................................................................................................... 8
   2.2 Specific Objectives ....................................................................................... 8
3.0       Changes to the Conservation Strategy since 2007 ..................................... 10
   3.1 Forestry Commission Changes since 2007 ................................................ 10
   3.2 Priority Species Designations ...................................................................... 11
   3.3 Butterfly Conservation Landscape Projects ................................................. 16
   3.4 Strategy Sites and Priority Site Grade changes .......................................... 16
4.0       Conservation Strategy Context.................................................................... 17
   4.1 Importance of woodlands for Lepidoptera .................................................. 17
   4.2 Changing status of Butterflies & Moths in woodlands and on the FC estate
              17
   4.3 Forestry Commission management for Butterflies & Moths to date ............ 19
   4.4 Current information on the status of Butterflies and Moths on FC land ..... 20
5.0       Prioritising and Grading Priority Sites .......................................................... 21
   5.1 Methodology ............................................................................................ 21
   5.2 Distribution of sites by grading ................................................................... 22
   5.3 Results of Strategy Review 2012................................................................. 23
   5.4 Summary Statistics 2012/13 ....................................................................... 27
6.0       References .................................................................................................. 29
7.0       Useful reading ............................................................................................. 30
Acknowledgements ................................................................................................ 31
Appendix 1: Species Grading ................................................................................. 32
Appendix 2. Lepidoptera found on Forestry Commission Land, their conservation
status and requirements......................................................................................... 35
Appendix 3. National Summary of Forest District Priority Sites by Designation ..... 40
Appendix 4: Key policies & strategies affecting Forestry and Biodiversity post-2010
............................................................................................................................... 41
Appendix 5i. Priority Forestry Commission Lepidoptera sites in England by FC
District 2012 ........................................................................................................... 42
Appendix 5ii. Priority Forestry Commission Lepidoptera sites in England by FC
District 2012 associated maps and tables .............................................................. 49
Central Forest District ............................................................................................ 51
East Forest District ................................................................................................. 54
North District .......................................................................................................... 58
South District .......................................................................................................... 62
Ampfield Wood ....................................................................................................... 62
West District ........................................................................................................... 68
Yorkshire District .................................................................................................... 74
Westonbirt Arboretum ............................................................................................ 78
Lepidoptera on Forestry Commission Land in England; Conservation Strategy 2007 - 2017 Review 2012/13
Foreword

Butterflies and moths are known to be valuable indicators of the changes affecting
the wider countryside and recent studies suggest that a large number of species
are in rapid decline. Butterflies and moths are one of the most threatened wildlife
groups that inhabit the Forestry Commission Estate and as active land managers;
we are able to play an important role in their future.

The Priority Lepidoptera Sites identified in this strategy cover approximately 2000 –
8000ha of woodland and associated open habitats such as heathland and
limestone pavement. These sites represent approximately 3% of the total Forestry
Commission England landholding and 56% of them have SSSI designation.

They support populations of some of the UK’s most endangered woodland
butterflies and moths including the Pearl-bordered Fritillary and Wood White, as
well as the Argent & Sable. The reasons behind the decline of these Biodiversity
Action Plan Priority Species are complex; however a lack of woodland
management is often cited as an important cause and one that we can act on to
support the conservation of these species.

Managing woodlands for Lepidoptera will make an important contribution to many
other work programmes taking place across the Forestry Commission England
Estate including work to enhance the condition of SSSI, Ancient Woodland
restoration and Biodiversity Action Plan habitat restoration. With the potential
demand for small diameter hardwoods on the increase through developments in
the wood fuel industry, new opportunities for the sustainable management of key
Lepidoptera habitats are being created.

Many Lepidoptera depend on a network of early succession habitats, such as
young coppice as well as open, sunny rides for their long-term survival. If we are
to prevent the loss of key woodland Lepidoptera from Forestry Commission
woodlands, we need to target woodland management practices more directly and
more urgently at the Priority Sites listed in this Conservation Strategy.

Much good work has already been achieved by Forestry Commission staff working
closely with Butterfly Conservation colleagues. A key objective of this strategy is to
further raise awareness of the important sites, to ensure that they feature as
priorities in both the Design Planning process and the daily management of our
woodlands. It also provides the opportunity to underpin a bid for funding to give
the Forestry Commission the best chance of achieving the aims of the Strategy by
2017.

It is vital that the Forestry Commission’s delivery of Priority Site conservation and
management is viewed as part of a much wider programme of delivery to ensure
that these diverse and beautiful insects are allowed to increase and thrive in our
English woodlands.

Simon Hodgson
Chief Executive Forestry Commission England
Lepidoptera on Forestry Commission Land in England; Conservation Strategy 2007 - 2017 Review 2012/13
Summary

This strategy supports the agreement between Forestry Commission England (FC)
and Butterfly Conservation to conserve and increase populations of butterflies and
moths by increasing the level and urgency of woodland management across sites
considered to be important for Lepidoptera.

It is aimed at the foresters, conservation managers and planners in each Forest
District who are responsible for planning and implementing the management
activities across our landholding. It provides details of each priority site as well as
more generic information on the habitat requirements of each of our most
threatened species.

With advice and input from Forest District staff and Butterfly Conservation 140 sites
known to be important for butterfly and moth species were identified and formed
the basis of the strategy in 2007. Since the writing of the Strategy in 2007 the
twelve former FC Districts have been condensed to 7 (see Section 3.1). The total
number of woodlands that this review is based on has now changed to 136 as four
sites originally included were disposed by the Forestry Commission in 2012.

Priority Site grading has changed for 11 sites due to colonisations and extinctions
of priority species; an additional 24 sites have either increased or decreased in
‘Total Species Score’ (see Section 3.4 & 5.3). These changes are due to additional
survey effort and knowledge of a species occurrence on a site specific basis. The
following summarises the site changes:

   85 (change from 77) sites have been graded A because they support species
    such as the Pearl-bordered Fritillary and Argent & Sable that depend
    specifically on woodland and require urgent targeted management.

   27 (change from 33) sites are graded B. These sites support a lesser variety of
    priority species or species that are not so reliant on urgent woodland
    management such as the Grizzled Skipper.

   24 (change from 30) sites are graded C. These sites support more generalist
    priority or species of conservation concern such as the White Admiral.

The Strategy ‘Site Grading’ and ‘Species Scores’ are based on a total of 59
Lepidoptera species of which 38 are currently designated as both Section 41 and
UKBAP Priority Species; 16 butterflies and 22 moths.

33% of sites in the Strategy (45 out of the 136) are or have been part of a Butterfly
Conservation led Landscape project since 2007.

The objectives are to conserve populations of all priority Lepidoptera species on
FC land through active woodland management. This can only be achieved through
raising awareness of the urgency and necessity of each management activity on
the ground.    This strategy aims to do that through promoting a better
understanding of the sites important for Lepidoptera and the species requirements
across FC England.
Lepidoptera on Forestry Commission Land in England; Conservation Strategy 2007 - 2017 Review 2012/13
It should become a key document used to inform all stages of the management
process from Forest Design Planning through to operational planning and
management activity in the woods.

The Conservation Strategy will be reviewed with each Forest District on a rolling
programme to ensure that it is kept alive and relevant throughout its duration. The
overall success will be determined by the status of priority Lepidoptera at priority
sites at the mid-term review date of 2012 and again in 2017.

In 2012 the information within this Strategy has been reviewed. In 2017 a review of
this Strategy and your knowledge of these sites will help us to re-assess Priority
Site information and demonstrate conservation success.
Lepidoptera on Forestry Commission Land in England; Conservation Strategy 2007 - 2017 Review 2012/13
1.0    Introduction

1.1 Purpose of the Report

None of the work needed to maintain priority Lepidoptera is new to the Forestry
Commission – most Priority Sites simply need more action, and urgently. A
significant proportion of Priority Sites are Ancient Semi-natural Woodland Sites
(ASNW) or Plantation on Ancient Woodland Sites (PAWS), already managed for
their biodiversity value or under a programme of restoration back to semi-natural
woodland composition. In addition, a number of sites are designated as Sites of
Special Scientific Interest (SSSI), many for their butterfly and moth interest and are
currently being targeted under PSA agreement to ensure that 95% of the SSSI
network on the public forest estate is maintained in favourable or recovering
condition, while over time increasing the area of SSSI in favourable condition.

To put the conservation of Lepidoptera in context, approximately 3% of the FC
estate has been identified as a priority site. These sites form an integral part of
sustainable woodland management and should be maintained with the same set of
planning and operational tools that are used across the whole estate. Taking an
integrated approach means that these sites are managed alongside the wider
estate, supporting and contributing to other objectives such as Ancient Woodland
restoration, SSSI management and Habitat Action Plan targets.

The Forestry Commission is currently facing many challenges. Woodland
management for conservation depends on the availability of suitable markets for
small diameter hardwoods such as coppice. The most important market for such
products, the St. Regis paper mill closed in early 2006. Other challenges include
the increasingly wide range of objectives that have to be reconciled whilst
competing for the available resources. Securing additional resources will greatly
reduce the challenge of improving woodland habitats for Lepidoptera.

Although they can be managed alongside other objectives, woodland Lepidoptera
are dependent on regular habitat management for their survival. If forest
management does not provide, for example, open habitat on a rotational basis, or
rides become too shaded or connectivity between areas of suitable habitat is lost,
populations can quickly become extinct. It is crucial that Priority Sites, known to be
valuable due to their important Lepidoptera assemblages, feature highly in both the
Forest Design Plans and the Operational Site Assessments that take place before
operations commence, so that opportunities to enhance their habitats are not
missed.

This strategy aims to reduce the chances of local extinction happening by raising
awareness of these sites, of the Lepidoptera they support and of the general
habitat requirements of each of the species across the FC Estate. It does not aim
to provide detailed, site specific management prescriptions for each Priority Site,
but encourages you to seek guidance from Butterfly Conservation staff if required.

Priority Site species information contained within this strategy was correct at the
time of writing in 2007. It was collated using Butterfly Conservation monitoring
data and information from the Forest District staff. This is a working document - use
Lepidoptera on Forestry Commission Land in England; Conservation Strategy 2007 - 2017 Review 2012/13
the Forest District tables to record new sightings or management activities that may
have benefited Lepidoptera. In 2017 a review of this Strategy and your knowledge
of these sites will help us to re-assess Priority Site information and demonstrate
conservation success.

In 2012 the information within this Strategy has been reviewed using national
butterfly and moth data, District Ecologist and Butterfly Conservation
Regional staff comments.

The publication of this strategy in 2007 coincided with the recent National Review
of Biodiversity Action Plan (BAP) Priority Species by the Joint Nature Conservation
Committee (JNCC). A significant number of butterfly and moth species have been
added to the priority list, due to their increasing rarity within the British countryside.
Many of these species occur on Forestry Commission land, again emphasising the
ongoing commitment needed to ensure the objectives of this Conservation
Strategy are achieved.

2.0      Aims of the Conservation Strategy

2.1      Overall Aim

     Through active forest management planned for butterflies and moths, FC aims
      to conserve populations of all BAP Lepidoptera species on the Forestry
      Commission England Estate.

The priorities are to:

            Prevent further extinctions
            Stabilise colonies and numbers
            Increase populations and range of the BAP Priority Species that require
             targeted woodland management

2.2      Specific Objectives

This Conservation Strategy will achieve the overall aim above through:

     Ensuring that more urgent action is carried out within the Priority Sites that are
      listed in this Strategy and that this management is integrated with other
      management activities such as PAWS restoration, timber harvesting (including
      wood-fuel initiatives), SSSI condition enhancement and Habitat Action Plan
      restoration targets.

     Raising awareness and promoting a better understanding of Priority Sites and
      species to inform and guide all stages of Priority Site management from Forest
      Design Planning through to Operational Site Assessment and activity in the
      woods.

     Increasing communication between FC staff and Butterfly Conservation staff
      and to ensure that expert advice is sought by both parties when appropriate.
Lepidoptera on Forestry Commission Land in England; Conservation Strategy 2007 - 2017 Review 2012/13
     Encouraging further survey effort for Lepidoptera within FC woodlands,
      especially Grade A Priority Sites, and ensuring that records of scarce and
      threatened species are forwarded to Butterfly Conservation.

     Ensuring increased monitoring of Priority Sites results in more informed and
      responsive actions being taken in terms of planning and management activity.

     Achievement of the above objectives will be subject to available resources, and
      it will be vital to use this Conservation Strategy to underpin and secure
      additional funding from other parties to achieve the aims and objectives of this
      strategy.

2.3      Further objectives (subject to additional funding)

     FC will co-operate with partners over planned re-introductions, using Butterfly
      Conservation guidelines. It will be the responsibility of partners to assess
      feasibility and conduct actual re-introductions. FC will work with partners to
      create and maintain the desired habitat conditions.

     In co-operation with partners Butterfly Conservation will identify the key
      outstanding research issues for the conservation of Woodland Lepidoptera and
      will discuss with research providers.

     Butterfly Conservation will produce, in co-operation with partners, an advisory
      leaflet to provide the best current information on woodland management for
      Lepidoptera.

      The success of this strategy will be determined by how fully the
      objectives of this Conservation Strategy have been implemented and met.
      This will be assessed by the number of sites in 2017 maintaining or
      improving their populations of Priority Lepidoptera and favourable habitat
      condition status.
3.0    Changes to the Conservation Strategy since 2007

3.1    Forestry Commission Changes since 2007

Appendix 4 details the key policies & strategies affecting Forestry and Biodiversity
post-2010

Forestry Commission Districts
Following the 2010 Government Spending Review a restructuring programme was
implemented across Forestry Commission England which resulted in the reduction
of administrative areas for the public forest estate from 11 to 6 Forest Districts plus
the National Arboretum at Westonbirt. Following this reorganisation each Forest
District now has an Ecologist responsible for providing a support role to the local
forest planning and operational teams and maintaining partnership working with
key conservation stakeholders. The District Ecologists are responsible for
championing the joint strategy for Lepidoptera on the public forest estate and
ensuring that its objectives are delivered via strategic and operational plans.
Please refer to Figure 1 which depicts the former and current Forest Districts.

Figure 1. Left: Forestry Commission Districts 2007 Right: Forestry Commission
Districts 2012

Site Disposals
Prior to the Governments formation of the Independent Panel on Forestry and the
associated moratorium on site disposals some four priority sites originally listed in
the 2007 strategy have been disposed of. These sites are listed in Table 1.
Sites Disposed
Priority Site                           FC District 2007-
Grade           Sites Disposed          2012                Sites Disposed species Asc.
A               Rewell Wood             South               Pearl-bordered Fritillary
                                                            White Admiral, White Letter
B               Bramfield Wood          East                Hairstreak
                Balls Wood and                              White Admiral, Silver-washed
C               Woodhouse               South & East        Fritillary
Totals                              4

Table 1. Disposed Priority Sites and their Priority Site Grade 2007-2012

3.2 Priority Species Designations

NERC Section 41 species and former UKBAP
Of the 59 species included in this Strategy when it was written, 28 were designated
as UKBAP Priority and 14 were under review. Following the UKBAP review in 2007
there were 38 UKBAP Priority Species included; 16 butterflies and 22 moths.

Following UK and International policy change, the approach to species and habitat
conservation in the UK is now being approached at a country level within the UK
Biodiversity Framework.

Consequently, after this Strategy was published all of the 38 UKBAP Priority
Species included were designated in England as Section 41* species listed in the
Natural Environment and Rural Communities Act 2006. This legislation requires
that public bodies, such as the Forestry Commission, have a duty under section 40
of the same act to “have regard” to the conservation of biodiversity in England
when carrying out their normal functions. Since the publication of the NERC Act
(2006) the Lepidoptera Strategy is of even greater strategic importance.

Biodiversity 2020 NERC Section 41 species Action targets
The Biodiversity 2020 Terrestrial Biodiversity Group was requested to highlight the
actions required to assist in the recovery of all species designated as S41. The
group, consisting of a range of expert species ecologists, identified and prioritised
a set of actions for each species.

The actions are considered to be those required for S41 species to recover as well
as meet Outcome 3 of the Biodiversity 2020 strategy written by the government.
The actions were updated in January 2014 and Table 2 has been adapted to
reflect the species included in the Lepidoptera Strategy. The full table can be
downloaded from
http://publications.naturalengland.org.uk/publication/4958719460769792.

All of the actions listed require the ‘Land and Woodland Management’ as
their delivery mechanism.
The definitions of ‘Priority Group’ are as follows:
1 = global concern
2= European concern
3=national concern
4=other national concern
*The definition of this designation is as follows “Species of principal importance for the purpose of conserving
         biodiversity” covered under section 41 (England) of the NERC Act (2006) and therefore need to be taken into
         consideration by a public body when performing any of its functions with a view to conserving biodiversity.

Section 41 and UKBAP Priority Species that require urgent targeted management:
                                                                                                                            Priority
                                               FC                                                                           Group (as
                       Taxon name (=
Common name                                    Species         Action text                           Action priority        defined by
                       synonym)
                                               Score                                                                        Natural
                                                                                                                            England)
                                                               EWGS designed to
                       Rheumaptera
Argent and Sable                                     15        encourage open woodland               Medium                      3
                       hastata
                                                               with abundant birch re-growth.
                                                               Agri-env schemes designed to
                                                               preserve its specialised early-
                                                               successional habitat. Its
Basil-thyme
                       Coleophora tricolor           15        specific requirements are not         Medium                      3
Case-bearer
                                                               fully understood & BC should
                                                               be consulted for detailed
                                                               advice.
                                                               EWGS to maintain the open
                                                               glades and clearings in
Betony Case-           Coleophora
                                                     15        woodlands and encourage               Medium                      3
bearer                 wockeella
                                                               growth of the larval food plant
                                                               (betony).
                                                               EWGS to maintain the open
Drab Looper            Minoa murinata                15        areas in ancient woodlands            Medium                      3
                                                               which this species requires.
                                                               Secure habitat management
                                                               at all sites through HLS where
                                                               possible/ practical; this
Duke of Burgundy       Hamearis lucina               15                                              Urgent                      3
                                                               species is covered under the
                                                               Farmland Butterfly Initiative
                                                               (FBI).
                                                               Secure habitat management
                                                               at all sites through EWGS
Heath Fritillary       Melitaea athalia              15                                              Medium                      3
                                                               where HLS not possible/
                                                               practical.
                                                               Secure habitat management
                                                               at all sites through HLS (FBI)
High Brown                                                     where possible/ practical; this
                       Argynnis adippe               15                                              Urgent                      3
Fritillary                                                     species is covered under the
                                                               Farmland Butterfly Initiative
                                                               (FBI)
                                                               EWGS to preserve the open
Lunar Yellow
                       Noctua orbona                 15        woodland glades where this            Medium                      3
Underwing
                                                               species occurs.
                                                               Secure habitat management
Pearl-bordered         Boloria                                 at all sites through EWGS
                                                     15                                              Urgent                      3
Fritillary             euphrosyne                              where HLS not possible/
                                                               practical.
                                                               EWGS (based on regular
Small Pearl-                                                   rotational ride/ glade
                       Boloria Selene                15                                              Medium                      3
bordered Fritillary                                            maintenance, or a suitable
                                                               coppicing regime).
                                                               EWGS (based on regular
                                                               rotational ride/ glade
Wood White             Leptidea sinapis              15                                              Medium                      3
                                                               maintenance, or a suitable
                                                               coppicing regime).
Section 41 and UKBAP Priority Species where woodland is a secondary habitat but active
management necessary:

                                                                                        Priority
                                                                                        Group
                            FC                                                          (as
Common       Taxon name                                                      Action
                            Species   Action text                                       defined
name         (= synonym)                                                     priority
                            Score                                                       by
                                                                                        Natural
                                                                                        England)

                                      Agri-env schemes to ensure suitable
                                      management of the hedgerows &
Barberry     Pareulype                woodland edges where it occurs; in
                              5                                              Medium        3
Carpet       berberata                particular hedgerows should be
                                      trimmed in late autumn, after larvae
                                      have pupated.
Barred                                EWGS to maintain/ restore wide
             Trichopteryx
Tooth-                        5       woodland glades and edge habitats      Low           3
             polycommata
striped                               where privet can flourish.
Buttoned     Hypena
                              5
Snout*       rostralis
                                      Agri-env schemes, tailored to
                                      manage hedgerows and woodland
                                      edges in a sympathetic way for this
Brown        Thecla                   species. Involves re-planting
                              5                                              Medium        3
Hairstreak   betulae                  hedgerows (with blackthorn
                                      included) as well as more
                                      appropriate management of existing
                                      hedgerows.
                                      EWGS to maintain woodland glades
Clay Fan-    Paracolax
                              5       and other suitable open areas in       Medium        3
Foot         tristalis
                                      woodlands.
                                      Species requirements not fully
                                      understood, but long-term rotational
Common       Pechipogo                coppicing & occasional felling of
                              5                                              Medium        3
Fan-foot     strigilata               mature trees (to provide gaps) helps
                                      & this should be provided through
                                      EWGS.
                                      Agri-env schemes designed to
                                      preserve the damp grasslands and
Dingy        Cyclophora
                              5       heathlands where the required          Medium        3
Mocha        pendularia
                                      willows grow inn un-shaded
                                      situations.
Dingy        Erynnis                  EWGS (based on regular rotational
                              5                                              Low           3
Skipper      tages                    maintenance of wide glades/rides).
                                      EWGS to provide the coppiced
False        Cyclophora
                              5       woodland & clearings/ glades it        Medium        4
Mocha        porata
                                      requires.
                                      Agri-env schemes to preserve its
             Hipparchia
Grayling                      5       specialised, early successional        Low           3
             semele
                                      habitat.
Grizzled     Pyrgus                   EWGS (based on regular rotational
                              5                                              Low           3
Skipper      malvae                   maintenance of wide glades/rides).
Priority
                                                                                           Group
                               FC                                                          (as
Common       Taxon name                                                         Action
                               Species   Action text                                       defined
name         (= synonym)                                                        priority
                               Score                                                       by
                                                                                           Natural
                                                                                           England)
                                         Agri-env schemes suitably tailored
                                         to maintain the specific (wet boggy
Large        Coenonympha
                                 5       mire) habitat, with water levels       Medium        2
Heath        tullia
                                         properly maintained and sites not
                                         overgrazed.
                                         Secure habitat management at all
                                         sites through HLS where possible/
Marsh        Euphydryas
                                 5       practical; this species is covered     Urgent        3
Fritillary   aurinia
                                         under the Farmland Butterfly
                                         Initiative (FBI)
                                         Agri-env schemes to ensure
Narrow-                                  abundant larval food plant & nectar
bordered                                 sources. N.B. often occurs in same
             Hemaris tityus      5                                              Medium        3
Bee Hawk-                                grasslands as marsh fritillary and
moth                                     prescriptions should accommodate
                                         both spp. where they both occur.
                                         Ensure that grazing of upland
Northern
             Aricia                      limestone grasslands is not too
Brown                            5                                              Medium        3
             artaxerxes                  intensive and thus accommodates
Argus
                                         the needs of this butterfly.
                                         EWGS designed to maintain
                                         suitable numbers of mature trees
Olive        Trisateles
                                 5       with damaged branches, producing       Medium        3
Crescent     emortualis
                                         the preferred larval food plant of
                                         withered leaves.
                                         Little-known species and a better
Scarce
                                         knowledge of its autecology is
Aspen        Sciota hostilis     5                                              Urgent        3
                                         required to fully understand the
Knot-horn
                                         reasons for its decline.
                                         EWGS to maintain suitable
Scarce
                                         woodland habitat with sufficient
Aspen        Phyllonorycter
                                 5       aspen; may need autecological          Medium        3
Midget       sagitella
                                         studies as well as reasons for
Moth
                                         decline may not be fully known.
Silver-
             Plebejus                    Agri-env schemes to preserve its
studded                          5                                              Medium        4
             argus                       early successional habitat.
Blue
             Coscinia
                                         Agri-env schemes to maintain/
Speckled     cribraria
                                 5       enhance the specific heathland         Medium        1
Footman      subsp.
                                         habitat which this species requires.
             bivittata
Square-
             Xestia
spotted                          5
             rhomboidea
Clay*
Striped      Shargacucullia              Agri-env schemes to ensure good
                                 5                                              Medium        3
Lychnis      lychnitis                   supply of larval food plant.
Waved        Hydrelia
                                 5
Carpet*      sylvata
Section 41 and UKBAP Priority Species associated with mature trees:

                                                                                          Priority
                                                                                          Group
               Taxon         FC                                                           (as
Common                                                                         Action
               name (=       Species   Action text                                        defined
name                                                                           priority
               synonym)      Score                                                        by
                                                                                          Natural
                                                                                          England)
Dark                                   EWGS designed to preserve suitable
               Catocala
Crimson                        3       numbers of large, mature oaks in the    Medium        1
               sponsa
Underwing                              species range & habitat.
                                       Agri-env schemes designed to
                                       preserve mature & veteran oaks in
Heart Moth     Dicycla oo      3                                               Medium        4
                                       hedgerows, field margins, commons,
                                       woodland edge etc.
Light                                  EWGS designed to preserve suitable
               Catocala
Crimson                        3       numbers of large, mature oaks in the    Medium        1
               promissa
Underwing                              species range & habitat.
Scarce
               Moma
Merveille du                   3
               alpium
Jour*
                                       EWGS (based on regular rotational
White          Limenitis
                               3       ride/ glade maintenance, with mature    Low           3
Admiral        camilla
                                       woodland retained).
                                       Action centres on maintaining
                                       suitable elm trees, controlling Dutch
White Letter   Satyrium w-             elm disease where it still occurs and
                               3                                               Medium        3
Hairstreak     album                   re-planting with disease-resistant
                                       elms. EWGS, where appropriate, is
                                       one suitable mechanism for this.
                                       EWGS designed to preserve English
White-
               Cosmia                  Elms in woodlands & woodland edge
Spotted                        3                                               Medium        4
               diffinis                (including protection against Dutch
Pinion
                                       Elm Disease).
Other key species occurring in woodland:
                                       EWGS designed to maintain/
                                       enhance the damp woodland habitat
The            Chortodes
                               1       where it grows (including               Medium        4
Concolorous    extrema
                                       encouraging growth of larval food
                                       plant).

Table 2. Section 41 species actions
3.3 Butterfly Conservation Landscape Projects

Butterfly Conservation has been targeting Lepidoptera conservation through
working in a number of sites within a network across a distinct landscape for a
number of years. This was in response to the results of metapopulation research
which highlighted the importance of reducing species isolation through targeting
management across a number of connected sites in a landscape (Bourn & Bulman,
2005; Hanski 1998).

A recent publication which highlights five woodland case studies at the landscape
scale includes a number of Forestry Commission sites across the UK (Ellis et al
2012).

The number of FC Strategy sites that are involved with or included in Butterfly
Conservation Landscape projects changes on an annual basis as projects end and
others begin. Currently 33% of sites included in the Strategy (45 out of the 136) are
or have been part of a Butterfly Conservation led landscape project since 2007.

3.4 Strategy Sites and Priority Site Grade changes

The total number of woodlands that this 2012 Strategy review is now based on has
decreased to 136 from 140 in 2007 as four sites originally included were disposed
of by the Forestry Commission in 2012.

Priority Site Grading has changed for 11 sites due to colonisations and extinctions
of priority species; an additional 24 sites have either increased or decreased their
Total Species Score with an average change in score of +/-10 (see Section 5.3).

Priority                 Original
Site       2012          Strategy        Post
Grade      Review        2007            Disposals
A                   85              77         76
B                   27              33         32
C                   24              30         28
Totals          136            140            136

Table 3. Total number of Priority Sites Grades 2007 and 2012
4.0      Conservation Strategy Context

4.1      Importance of woodlands for Lepidoptera
     Woodland is an extremely important habitat for British Lepidoptera. Sixteen
      species of butterfly (out of Britain’s 59 resident species) rely on woodland in all
      or part of their range or have a large proportion of their population associated
      with woodlands (Clark et al 2011).
     Nearly two thirds of all British butterflies will use woodland as breeding habitat.
      (Fox et al 2011).
     There are around 2500 species of moths in Britain. Moths are found in almost
      all habitats and occupy a variety of niches but around 580 species of the larger
      British moths occur regularly in woodland habitat from open clearings to ancient
      woodland (Clark et al 2011).
     Open space within woodland provides a refuge for species of open habitats.
      The FC Estate includes important non-woodland habitats, including unimproved
      grasslands, upland and lowland heath and mires.

4.2      Changing status of Butterflies & Moths in woodlands and on the FC estate

During the 20th Century there has been an increasingly rapid decline in woodland
butterflies with many species disappearing from much of their former range.
Recent data analysis using the UK Butterfly Monitoring Scheme highlights that
specialist species that occur in woodlands (16) have declined by 56% since 1990
(Clarke et al 2011). It suggests that butterflies associated with woodland clearings
have undergone a marked decline of 80% since 1985.

In the recent publication The State of the UK’s Butterflies 2011, the 10 year
indicator trend using UKBMS data from 1999-2009 suggests that all butterflies
occurring in woodland have declined by 51%.. Butterflies associated with active
woodland management such as coppice clearings and ride management continue
to suffer declines including the Duke of Burgundy and Pearl-bordered Fritillary
which have declines in their 10 year population trend by 46% and 42% respectively
(Fox et al. 2011).

Trends of species that are associated with mature woodland like the Speckled
Wood and Silver-washed Fritillary have increased their populations by 42% and
38% respectively. This suggests that woodland conditions have become
increasingly shaded with a dense canopy, conditions under which these shade-
tollerant species can do well.

Moths have also seen a decline. The number of moths caught in Rothamsted light
traps has decreased by 28% over a 40 year period between 1968 to 2007. Since
the last report (Fox et al. 2006) the UKBAP list of moths increased to 81 and an
additional 71 widespread but rapidly declining larger moths were included.
However following the UKBAP review 142 moth species are now listed as Species
of Principal Importance in England under section 41 of the NERC Act 2006 (Fox et
al 2013).
Figure 2.   Graphs displaying the trends in woodland butterflies from 1990-
2009

Butterflies and moths are a well-studied group and the decline of so many
woodland Lepidoptera may indicate a serious, yet less well-documented, threat to
other woodland fauna and flora. A paper by Thomas et al. (2004) showed that
butterflies have declined by 71% since the 1970s, substantially more than the
decline recorded for British birds (54% over 20 years) and plants (28% over 40
years).

The overall decline in Lepidoptera has been strongly linked to changing
management, structure and composition of woods, e.g. increasing shade, fewer
open spaces and loss of plant diversity. Woodland management has altered
rapidly, with a change from extensive coppicing in the late 19 th Century in favour of
high forest and the widespread introduction of conifers to previously broad-leaved
woodland. Many of the woodlands that were not planted with conifers have
undergone 100 years of neglect, especially those in private ownership.

The darker conditions that accompany maturing conifer plantations are unsuitable
for most BAP Priority Lepidoptera. Few woodland butterflies can cope with well-
shaded habitat and this has led to a decline in the majority of species that depend
on early succession. Only the Speckled Wood, a species that can tolerate shaded
conditions more than most other butterflies has increased in abundance. Conifer
feeding moths, such as the Spruce Carpet, Pine Beauty and Satin Beauty have
also done well (Fox et al. 2006b) although the Bordered White, another conifer
specialist has declined (-33% in 35 years).

The impact of increasing deer numbers on ground flora has been identified as a
further threat to Lepidoptera. In some circumstances they can prevent wood re-
growth (thus affecting ground flora), the quality of coppice re-growth and,
occasionally, may directly graze out food plants. The White Admiral has seen a
50% decline in numbers, thought to be due to Honeysuckle being browsed out by
deer (Feber et al. 2001).

The intensification of agriculture and removal of hedgerow and woodland
fragments have left much woodland isolated, reducing the chances of re-
colonisation after local extinctions. The remaining woods are important as refuges
for grassland as well as woodland species. This increases the need to maintain
habitat variability within each individual woodland block, where habitat
fragmentation in the surrounding countryside is high.

The Forestry Commission was set up in 1919 with the objective of planting enough
forest to provide a strategic reserve of timber for future wars or national
emergencies. Afforestation by the Forestry Commission took place at an
unprecedented scale following the First World War, often with minimal regard for
existing habitats such as native woodland. This undoubtedly had a negative
impact on our woodland Lepidoptera.

However, many PAWS still retain some of the rich flora of their past and have very
good potential to support sustainable populations of Priority Lepidoptera species.
Over recent years, the widespread implementation of PAWS restoration through
the gradual removal of conifer and the use of natural regeneration to create broad-
leaved semi-natural woodland has given rise to more species rich woodlands.
However the move towards continuous cover forestry may have an adverse impact
on Lepidoptera populations reliant on the early successional stages once provided
through clearfelling. It is important that plentiful, well-connected open spaces such
as rides are available within these woodlands to ensure that adequate habitat is
available for Lepidoptera.

In addition FC are now taking a landscape-scale approach to habitat re-creation,
and restoration projects are resulting in the creation of larger and more sustainable
semi-natural habitat mosaics. FC woods are now representing a vital conservation
resource for butterflies and moths in England.

4.3   Forestry Commission management for Butterflies & Moths to date

FC management for butterflies dates back to the 1970s, with practice evolving over
the years. Management has developed from small-scale work in limited reserve
areas to whole-wood management which provides greater, and more sustainable,
conservation opportunities.

Small-scale management, using hand or mechanised hand tools in places like
Bernwood proved unable to slow succession sufficiently to provide habitat for key
early successional species. Larger-scale management plans developed in the late
1970s and early 1980s at places such as Bernwood and Haugh Wood created
large open spaces, in particular ‘box junctions’ which could be managed by
machine.

At the same time ride management suitable for broad, mechanised application was
developed by Forest Research and widely applied. However, these practices did
not live up to their initial promise because natural succession to grasses and other
coarse vegetation meant that managed edges and glades lacked a bare
ground/woodland herb component and became unsuitable for early succession
species. Considerable effort has been made to overcome this problem, including
the use of forage harvesters to remove cut vegetation and selective herbicide
application. This has not produced habitat comparable to coppice or high forest
felling, although has resulted in suitable habitat in some situations.

Clear fells and windblow particularly of conifer, have provided valuable early
succession habitat, for example in South East England, and in the wildlife corridors
developed largely by felling middle-aged conifer in Wyre Forest. The long-term
suitability of these sites will depend on follow-up management, but initial progress
has been made.

Butterfly management on a ‘whole wood’ basis is now being developed linked to
both SSSI and Forest Design Plans (FDPs). In places like Haugh Wood, Wyre
Forest, Whitbarrow, Chiddingfold and Bernwood, management plans have
incorporated the needs of key species of Lepidoptera and implemented actions to
gain positive outcomes. FDPs and SSSI plans in the future must attempt to act on
this landscape-scale to halt the loss of BAP Priority Lepidoptera Species. There is
an important role for Butterfly Conservation in promoting work across land
ownership boundaries to create and maintain linkages.

In February 2006 the St. Regis paper mill closed. This provided by far the most
important market for small diameter hardwoods including coppice. Its closure
makes the sourcing of new and reliable markets vital if affordable and sustainable
management is to be achievable into the future.

4.4 Current information on the status of Butterflies and Moths on FC land

The Species Action Plan for Butterflies on Forestry Commission Land was
published in 2000. The butterfly species information contained in that plan was
updated for the Conservation Strategy and information on moths was included.

The list for moths is still not exhaustive; however, further survey effort for moths
within FC woodlands is to be encouraged and records of scarce and threatened
species should be forwarded to Butterfly Conservation.

Following the UKBAP Review and NERC Act S41 designations a review of the
actions required was published and the actions involving FC are detailed in
this review (Section 3.2).
5.0     Prioritising and Grading Priority Sites

5.1     Methodology

Site Grading

All Priority Sites identified within this strategy review are important and collectively
represent the best sites for Lepidoptera across the Forestry Commission England
estate. In 2007 and for this review in 2012, sites have been graded using the
National Moth Recording Scheme and Butterfly Monitoring Scheme records*, a
further grading review will be required in 2017 to assess that efforts have been
directed at those sites which support the most vulnerable species and which would
have been most affected through lack of management.

Sites have been graded A, B or C depending on:

     The status of the species (BAP or Species of Conservation Concern)
     The degree to which the individual species depend on suitably managed
      woodland habitats for their survival.
     The number of species present.
     The combined species score for an individual site**

Grade       Description                                                   Species
                                                                          Score **
Grade A     Grade A sites are the highest priority sites. They support    >15
            at least one woodland BAP Priority Species that
            depends on targeted and specific woodland
            management operations such as ride management or
            coppicing.
Grade B     Grade B sites support more generalist priority or species     6-15
            of conservation concern that are not so dependent on
            woodland habitats or management to maintain early
            successional stages.

Grade C     Grade C sites also support more generalist priority or
Species Grading

Appendix 1 details the NERC Section 41 and former Priority BAP Species and
Species of Conservation Concern recorded throughout the FC England Estate as
well as associated species scores used to Grade sites A, B or C.

Habitat Condition Assessment

The habitat condition score describes how suitable the habitat is for the specific
Lepidoptera present upon the site and how well the site is being managed for these
species. The habitat condition of most sites has not been assessed in this review
and will be recommended for the full review in 2017.

Forest District staff were responsible for providing the assessment in 2007 based
on advice from Butterfly Conservation as to the requirements of the species. The
condition assessments used are as follows:

Habitat            Description
Condition
Favourable         The site is already in favourable condition and current ongoing
Improving          management efforts could only work to improve the site further.
Favourable         The site is being managed appropriately for the Lepidoptera
                   species present.
Unfavourable       Through appropriate management the site is beginning or is
Recovering         predicted to show a move towards favourable condition,
                   however it has not yet reached favourable condition.
Unfavourable       The site is in unfavourable condition but is neither declining nor
No Change          recovering. Further management is required to achieve
                   unfavourable recovering status.
Unfavourable       The site has not been appropriately managed for Lepidoptera
Declining          for sometime or has undergone a damaging activity. Recovery
                   is possible and may occur if suitable management input is
                   made.

5.2   Distribution of sites by grading

The map and tables in Appendix 5 list the Priority Sites included in this Strategy
that were reviewed in 2012 together with the Grade and habitat condition of the
Site following the results. This information has been mapped on GIS and is
available on request. The data is held by the England Biodiversity Information
Officer to whom any updated information should also be sent.

The list of sites is not exhaustive, if key species are found on FC land that
are not listed here, Forest Districts are encouraged to consult with Butterfly
Conservation over management issues if necessary.
5.3 Results of Strategy Review 2012

Site Specific Priority Grading

Priority Site Grading has changed for 11 sites due to colonisations and extinctions
of priority species; an additional 24 sites have either increased or decreased their
Total Species Score with an average change in score of +/-10.

Of the 11 Priority Site Grade changes, 10 have been positive with 9 sites now
graded as A, where 5 woods were regraded from B to A and 4 from C to A, the
remaining positive change was from C to B. Of these 10 sites half are or have
been involved in a Butterfly Conservation Landscape project. The sites have been
re-graded due to a change in Lepidoptera interest, with 80% of sites been
colonised by a single species since 2007 and recorded in 2012 i.e. Dingy Mocha
(2) Drab Looper (3), Small Pearl-bordered Fritillary (1) and Pearl-bordered Fritillary
(2) (see Table 4.).

There have been two sites (20%) with a Priority Site Grade change from B to A
which have had more than one additional species recorded since 2007. The site
with the biggest ‘Species Score’ change of 116 is Chickerell Woods in the East
District were 7 new moth and 6 new butterfly species have been recorded. The
other site with a ‘Species Score’ change of 45 is Vernditch in the South District
which has had the following additional three species recorded since 2007; Drab
Looper, Duke of Burgundy and Pearl-bordered Fritillary.

Red Lodge Wood, West District is the only site that has decreased in its grade with
a change from B to C due to Brown Hairstreak and White Letter Hairstreak not
recorded since 2009 and 2010 respectively.

Priority Species Colonisations and extinctions

Table 5 details the sites and additional species recorded since 2007 which has
increased their ‘Total Species Score’ but not affected the overall Priority Site
Grade. 58% of these sites are currently or have been involved in a Butterfly
Conservation Landscape project. 70% of the sites with an increase in the ‘Total
Species Score’ has been due to a single species colonisation and recorded in
2012. The other 30% of sites have had from two to four additional species
recorded in 2012. The site which has had the most new species recorded is the
Forest of Dean (Highmeadow) in the West District.

The following tables detail the 11 Priority Site Grade changes and the change in
the ‘Total Species Score’ on an additional 24 sites of which 17 have increased and
7 decreased but did not result in a grade change (see Section 3 & 5).
Change     Species       Additional
                   FC          Butterfly        Site     Site
                                                                  in Total   Not           Species
Site Name          District    Conservation     Grade    Grade
                                                                  Species    Recorded      Recorded
                   2012        Project Name     2007     2012
                                                                   Score     2012          Since 2007
                                                                             White
Chicksands                                                                   Letter        X7 moths, x6
Woods              East                            B        A        116     Hairstreak    butterflies
                                                                                           Drab Looper,
                                                                                           Pearl-
                                                                                           bordered
                               South East                                                  Fritillary, Duke
Vernditch          South       Woods               B        A        45                    of Burgundy
                               Dukes on the
Charlton           South       Edge                B        A        15                    Drab Looper
                               South East
Eartham Woods      South       Woods               B        A        15                    Drab Looper
                                                                                           Pearl-
                               South East                                                  bordered
Houghton Wood      South       Woods               C        A        15                    Fritillary
                                                                                           Pearl-
                               Midlands                                                    bordered
Ribbesford Wood    West        Fritillary          C        A        15                    Fritillary
Tidenham Chase
Woods              West                            C        A        15                    Drab Looper
                                                                                           Small Pearl-
Cann Wood, Plym                                                                            bordered
Forest             West                            C        A        15                    Fritillary
Moreton            South                           B        A         5                    Dingy Mocha
Park & Queen
Copse              South                           C        B         5                    Dingy Mocha
                                                                             White
                                                                             Letter
                                                                             Hairstreak,
                                                                             Brown
Red Lodge Wood     West                            B        C         -8     Hairstreak
  Chicksands       Pearl-bordered Fritillary, Small Pearl-bordered Fritillary, Wood White, Dingy Skipper,
Woods-Additional   Brown Hairstreak, Purple Emperor, Argent & Sable, Common Fan-foot, Clay Fan-foot,
   Species=               White-line Snout, Waved Carpet, Drab Looper, Coleophora wockeella

     Table 4. Priority Site Grade Changes
Change
                   FC          Butterfly       Site    Site               Species Not    Additional Species
                                                               in Total
Site Name          District    Conservation    Grade   Grade              Recorded       Recorded Since
                                                               Species
                   2012        Project Name    2007    2012               2012           2007
                                                                Score
                                                                                         Pearl-bordered
                                                                                         Fritillary, Drab
                               Herefordshire                                             Looper, Fletchers
Forest of Dean                 Wood White                                                Pug, Common Fan-
(Highmeadow)       West        Project             A       A   36                        foot
                                                                                         Small Pearl-
                                                                                         bordered
                               Saving Devons                                             Fritillary,White
Great                          Precious                                   Grizzled       Admiral,Silver-
Plantation         West        Pearls              A       A   19         Skipper        washed Fritillary
Somerford                                                                 Dark Green     Duke of Burgundy,
Common             West                            A       A   19         Fritillary     Marsh Fritillary
                               Dukes on the                                              Pearl-bordered
Grovely Wood       South       Edge                A       A   15                        Fritillary
                                                                                         Possible Pearl-
Whiteley                                                                                 bordered Fritillary
Pastures                                                                                 colonisastion or
(formally Botley               South East                                                unauthourised
Wood)              South       Woods           A       A       15                        introduction
Cannock                        Still being
Chase              Central     assessed            A       A   15                        Argent and Sable
Stonedown          South                           A       A   15                        Drab Looper
                               Herefordshire
                               Wood White                                 White Letter
Wigmore Rolls      West        Project             A       A   15         Hairstreak     Drab Looper
                                                                          Dark Green
                                                                          Fritillary,
                                                                          Silver-
                                                                          washed
                                                                          Fritillary,
                                                                          ?Dingy
                               Saving Devons                              Skipper,
                               Precious                                   Grizzled       Small Pearl-
North Wood         West        Pearls              A       A   13         Skipper        bordered Fritillary
                               Herefordshire                                             Barred Tooth-
                               Wood White                                 Argent and     striped, Fletchers
Dymock Woods       West        Project             A       A   6          Sable          Pug, Wood White
Friston Forest     East        (part) HLS          A       A   5                         Olive Crsent
Wareham
Forest             South                           A       A   5                         Dingy Mocha
Forest of Dean                                                                           Pearl-bordered
(Drybrook Rd                                                                             Fritillary ,
Station etc)       West                            A       A   5          Grayling       Dichomeris ustellala
Bedford                                                                   Square-        White Letter
Purlieus           Central     WREN; BIFFA         B       B   3          spotted Clay   Hairstreak
                               North York
                               Moore Dukes
Dalby Forest       Yorkshire   and Pearls          A       A
Waverley Wood
& Weston                       Midlands                                   Grizzled       Silver-washed
Wood               West        Fritillary          B       B   1          Skipper        Fritillary
                                                                                         Schiffermullerina
Shrawley Wood      West                            B       B   1                         grandis

         Table 5. Priority Site Increases in Total Species Score
Additional
                FC         Butterfly             Site    Site    Change                                Species
                                                                            Species Not
Site Name       District   Conservation          Grade   Grade   in Total                              Recorded
                                                                            Recorded 2012
                2012       Project Name          2007    2012    Species                               Since
                                                                  Score                                2007
                                                                            Argent and Sable,
                                                                            Pearl-bordered
                                                                            Fritillary, Small Pearl-
Chiddingfold    South      South East Woods        A      A      -45        bordered Fritillary
Parkhurst                                                                   Pearl-bordered
Forest          South      South East Woods        A      A      -15        Fritillary
Blackdown                                                                   Brown Hairstreak,
Forest          West                               A      A      -10        Marsh Fritillary
Lydford         West                               A      A      -5         Grizzled Skipper
                           Herefordshire Wood
Oversley Wood   West       White Project           A      A      -2         Grizzled Skipper
Mortimer                   Herefordshire Wood
Forest          West       White Project           A      A      -1         Dark Green Fritillary
                           Conserving the Wood                              Silver-washed
Purslow Wood    West       White butterfly         A      A      -1         Fritillary

      Table 6. Priority Site Decreases in Total Species Score
5.4      Summary Statistics 2012/13

                                               Total number
                       Grade   Grade   Grade      of sites
  Forest                                       assessed as
               Total     A       B       C
  District                                     “Favourable”
                       Woods   Woods   Woods
                                                  habitat
                                                 condition    BAP Priority Species that are principally woodland species and require urgent targeted
                                                   2007                                            management:

  Central       21      13       4       4          7
                                                              Argent & Sable, Wood White, Small Pearl-bordered Fritillary

      East      34      16      10       8         15         Basil Thyme Case-bearer, Lunar Yellow Underwing, Duke of Burgundy, Heath Fritillary,
                                                              Pearl-bordered Fritillary, Small Pearl-bordered Fritillary, Wood White

      North     12       8       0       4          5         Duke of Burgundy, High brown fritillary, Pearl-bordered Fritillary, Small Pearl-bordered
                                                              Fritillary

  South         27      15       8       4         15         Argent & Sable, Coleophera wockeella, Drab Looper, Duke of Burgundy, Pearl-
                                                              bordered Fritillary, Small Pearl-bordered Fritillary, Wood White

      West      34      26       4       4         16         Drab Looper, Duke of Burgundy, Heath Fritillary, Pearl-bordered Fritillary, Small Pearl-
                                                              bordered Fritillary, Wood White

Westonbrit      1        0       1       0          0

 Yorkshire      7        7       0       0          3
                                                              Argent & Sable, Duke of Burgundy, Small Pearl-bordered Fritillary
  TOTAL        136      85      27      24         61
6.0   References

Bourn N A D, Bulman C R (2005) Landscape scale conservation, theory into
practice. In: Kuhn E, Feldmann R, Thomas J A, Settele J (eds) Studies on the
Ecology and Conservation of Butterflies in Europe. Vol 1: General Concepts and
Case Studies, pp111-112, Butterfly Conservation, Wareham, Dorset

Clarke, S. A., Green, D. G., Bourn, N. A. & Hoare, D. J. 2011 Woodland
Management for butterflies and moths: a best practice guide. Butterfly
Conservation, Wareham.

Defra. 2006. Working with the grain of nature - taking it forward: volume I & II. Full
report on progress under the England Biodiversity Strategy 2002 - 2006. Defra,
London.

Ellis, S. Bourn, N. A. D. And Bulman, C. R. 2012 Landscape-scale conservation for
butterflies and moths: lessons from the UK. Butterfly Conservation, Wareham,
Dorset.

Feber, R.E., Brereton, T.M., Warren, M.S. & Oates, M. 2001. The impact of deer on
woodland butterflies: the good, the bad and the complex. Forestry. Vol 74. No 3,
pg 271 – 276.

Fox, R., Asher, J., Brereton, T., Roy, D. & Warren, M.S. 2006a. The State of
Butterflies in Britain and Ireland, Pisces Publications, Oxford.

Fox, R., Conrad, K.F., Parsons, M.S., Warren, M.S & Woiwod, I.P. 2006b. The
State of Britain’s larger moths. Butterfly Conservation and Rothamsted Research,
Wareham.

Fox, R., Parsons, M. S., Chapman, J. W., Woiwood, I. P., Warren, M. S. & Brooks,
D. R. 2013 The State of Britains Larger Moths 2013. Butterfly Conservation and
Rothemsted Research, Wareham, Dorset

Fox, R., Brereton, T. M., Asher, J., Botham, M, S., Middlebrook, I., Roy, D. B> and
Warren, M. S. 2011 The State of the UK’s Butterflies 2011. Butterfly Conservation
and the Centre for Ecology & Hydrology, Wareham, Dorset

Green, D.G. 2000. The Status of Lepidoptera in the New Forest: A report for the
Forestry Commission by Butterfly Conservation. Butterfly Conservation, Wareham.

Hanski 1998 Metapopulation Dynamics. Nature. Vol 396. Pg41-49

Liley, D., Brereton, T & Roy, D. 2004. The Current Level of Butterfly Monitoring in
UK Woodlands: And potential use of the data as a Biodiversity indicator to Inform
Sustainable Forestry. Butterfly Conservation Report to the Forestry Commission.
Butterfly Conservation Report S04-35, Wareham.

Thomas, J. Telfer, M.G., Roy, D.R., Preston, C.D., Greenwood, J.J.D., Asher, J.,
Fox, R., Clarke, R.T. & Lawton, J.H. 2004. Comparative Losses of British
Butterflies, Birds and Plants and the Global Extinction Crisis. Science. Vol 303. pp
1879-1881.

Appendix 4 only:
DEFRA (2011) Biodiversity 2020: A Strategy for England’s Wildlife and Ecosystem
Services. DEFRA, London, 2011.

DEFRA (2013) Government Forestry and Woodlands Policy Statement –
Incorporating the Government’s Response to the Independent Panel on Forestry’s
Final Report. DEFRA, January 2013.

Forestry Commission (2011) The United Kingdom Forestry Standard – The
Governments’ Approach to Sustainable Forestry. Forestry Commission, Edinburgh,
2011.

Forestry Commission (2011) Forests and Biodiversity – UK Forestry Standard
Guidelines. Forestry Commission, Edinburgh, 2011.

HMSO (2011) The Natural Choice: Securing the Value of Nature. DEFRA, 2011.

Independent Panel on Forestry (2012) Independent Panel on Forestry – Final
Report. Independent Panel on Forestry, July 2012.

Lawton, J.H. et al. (2010) Making Space for Nature: a Review of England’s Wildlife
Sites and Ecological Networks. Report to DEFRA. September 2010.

7.0   Useful reading

Specific species reports for certain areas are also available and can be requested
from Butterfly Conservation please see examples from the list below:

Asher, J., Warren, M.S., Fox, R., Harding, P., Jeffcoate, G & Jeffcoate, S. 2001.
The Millennium Atlas of Butterflies in Britain and Ireland, Oxford University Press,
Oxford.

Bulman, C. 2007. Woodlands – a vital habitat for butterflies and moths. Quarterly
Journal of Forestry. Vol 101. pp 29-39

Defra Factsheets are available for most BAP Priority Lepidoptera species with
habitat management recommendations included. These are available from Butterfly
Conservation or can be downloaded from the website www.butterfly-
conservation.org

Ellis, S. 2005. Conservation of the Duke of Burgundy (Hamearis lucina) butterfly on
the North York Moors 2005. Butterfly Conservation Report S05-37, Wareham.

Ellis, S. 2005. Conservation of the Pearl-Bordered Fritillary (Boloria euphrosyne)
butterfly on the North York Moors 2005. Butterfly Conservation Report S05-36,
Wareham.
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